Philonthus cribriventris Bernhauer, 1912: 43 Bernhauer & Schubert, 1914: 334 Bruch, 1915: 444 Blackwelder, 1944: 132 Herman, 2001b: 2793 Philonthus catamarcanus var. densior Bernhauer, 1916: 36 Scheerpeltz, 1933: 1335 de Maus, 2009: 39 Revision of the southern South American species of Philonthus Stephens (Coleoptera: Staphylinidae) 2595 Chani-Posse, Mariana Zootaxa 2010 2010-08-31 2595 1 70 779LY Bernhauer, 1912 Bernhauer 1912 [151,652,151,177] Insecta Staphylinidae Philonthus Animalia Coleoptera 24 25 Arthropoda species cribriventris      Philonthus cribriventris Bernhauer, 1912: 43;  Bernhauer & Schubert, 1914: 334(catalog);  Bruch, 1915: 444(catalog);  Blackwelder, 1944: 132(list);  Herman, 2001b: 2793(catalog); Chani-Posse, 2004: 230 (list).     Philonthus catamarcanus var. densior Bernhauer, 1916: 36;  Scheerpeltz, 1933: 1335(catalog); Chani-Posse  de Maus, 2009: 39(syn.)   Diagnosis.  Philonthus cribriventrismay be identified by the eyes shorter than temples, the basal area of abdominal terga with coarse punctuation and the transverse distance between punctures distinctly smaller than diameters of punctures, and the apex of median lobe subrectangular in dorsal view. It differs from other southern South American species of  Philonthusby the first segment of hind tarsus longer than last segment, and the apex of tergum 10 subacute in both sexes.   Redescription.Length of the body 6.2–7.0 mm. Coloration. Head black; thorax castaneous-piceous; elytra and abdomen piceous-black; antennae, palpi and legs brunneous. Head slightly wider than long (HW/HL= 1.10–1.05) and at basal third narrower than at distal third, distinctly to moderately narrower than pronotum (HW/PW= 0.80–0.75); dorsal surface coarsely and moderately punctate, with more than three postocular punctures and three to five epicranial punctures; distance separating medial interocular punctures on frons as large as distance separating medial punctures from lateral punctures; infraorbital ridge present, not extending beyond postgenal ridge. Eyes moderately shorter than temples (EL/TL= 0.80) seen from above (Fig. 3). Antennae with segment 1 distinctly shorter than segments 2 and 3 combined, segments 2 and 3 subequal in length, 4 and 5 elongate, 8 quadrate, 9–10 transverse. Labial palpus with last segment about 1.4–1.8 times as long as preceding segment. Maxillary palpus with last segment fusiform, about 1.5 times longer than segment 3. Submentum 1.5 times as long as mentum. Neck no more than 0.6 times as wide as head at widest point (NW/HW= 0.6–0.5). Pronotum slightly longer than wide (PW/PL= 0.9), distinctly narrowed anteriad; dorsal surface of pronotum with two rows of punctures, each with five to six punctures. Hypomeron not visible in lateral view. Basisternum with medial longitudinal carina moderately developed, bulbous. Metasternum with circummesocoxal ridge broadly rounded medially. Elytra at sides distinctly longer than pronotum at midline (EtL/ PL=1.6–1.5), about 1.5 times as long as elytra along suture (EtL/Etl= 1.7), and longer than wide at base of elytra (EtL/EtW=1.2–1.1); transverse distance between punctures about as large as diameters of punctures. Front tarsi as long as front tibiae; middle tarsi shorter than middle tibiae; first segment of hind tarsus as long as segments 2 and 3 combined, and longer than last segment (S1/S5= 1.5). Abdominal terga coarsely punctuate at basal area, distance separating punctures distinctly smaller than diameters of punctures; terga 2 and 3 with elevated area between basal lines punctuate, posterior basal line straight medially. Male sternum 7 with apical margin slightly notched at middle. Tergum 8 inboth sexes with posterior margin truncate. Male genitalia. Sternum 8 deeply emarginate medio-apically. Tergum 10 subacute medio-apically ( Fig. 50), with two long subapical macrosetae and 8–10 apical setae. Sternum 9 moderately emarginate apically with two long subapical macrosetae and one apical setae at each side ( Fig. 49). Aedeagus with median lobe gradually narrowed from base of paramere; apex of median lobe subrectangular (in dorsal view); paramere reduced and fused to median lobe ( Figs. 51–52). Internal sac with distinctly sclerotized structures ( Fig. 52). Female genitalia. Tergum 10 similar to that of male. Second gonocoxite with 2–5 strong setae along its outer margin, and stylus with two apical setae ( Fig. 53).  Bionomics.Unknown.  Geographical distribution.  P. cribriventrisis only known from a few localities in northwestern Argentina( Saltaand Tucumán) ( Fig. 147).   Type material. Lectotypeof  Philonthus cribriventris: ♂, with labels: “Rep. Argentina/ Prov. Tucuman/ 30.I.1908/ C. Bruch”, “  cribriventrisBrnh/ Typus”, “Chicago NHMus. / M. Bernhauer Collection”, “ Lectotype  Philonthus cribriventris Bernhauer, 1912/ Des. Chani-Posse de Maus 2008” ( FMNH).  Paralectotype: ♂, with labels: “Rep. Argentina/ Prov. Tucuman/ 30.I.1908”, “  cribriventrisBrnh/ Cotypus”, “Chicago NHMus. / M. Bernhauer Collection”, “ Paralectotype  Philonthus cribriventris Bernhauer, 1912/ Des. Chani-Posse de Maus 2008” ( FMNH).    Holotypeof  Philonthus catamarcanusvar. densior: ♀, “Rep. Argentina/ Prov. Tucuman/ 19 / Richter”, “ v. densiorBernh./ Typusunicus”, “ ChicagoNHMus. / Bernhauer Collection” ( FMNH).  Additional material: 19 specimens( 6 ♂, 4 ♀, 9 unsexed)   ARGENTINA: Tucumán:“Rep. Argentina / Prov. Tucumán / 19 / Richter”, “in   Bernhauerhandwriting " longicornis Gr." / det. Bernhauer”, “ v. densior Bernh./ det. Bernhauer”, “Chicago NHMus. / Bernhauer Collection” 1 ♂( FMNH);  Siambon,  VII. 1932, Bosq., 1 ♂, 1 ♀( MACN);  Anfama,  VII. 1933,  Philonthus cribriventrisBrnh./ 8, in Bernhauerhandwriting "  cribriventrisdet. Bernhauer Bosq. Don. Bruch", ChicagoNHMus Bernhauer Collection, 1 ♂, 1 ♀( FMNH),  1 specimen( MACN),  2 ♂, 1 ♀, 2 specimens( MLPA);  Trancas, Tacanas,  24.VIII.1973, L. Stange, 1 ♀( IMLA);  Richter, in green "cotypus", Bernhauerdeterm, in Bernhauerhandwriting "  cribriventrisBrh.", 2561  Philonthus cribriventrisBernh, 3 specimens( MLPA);   27.I.1908, C. Bruch, 2 specimens( MACN),  1 specimen( MLPA).   Salta: Dto. Victoria, Los Toldos, Rio Lipco, 8.18.  XII.1973, A. Sobral, 1 ♂( IMLA). FMNH Argentina Richter Chicago Typus 25 26 1 1 Tucuman holotype FMNH Bernhauer Bernhauer 25 26 1 1 [718,1266,375,402] 1932-07 MACN Bosq. Siambon 25 26 2 1 1 1933-07 FMNH Bernhauer & Chicago & Bernhauer Collection Bernhauer Bosq. Don. Bruch Anfama 25 26 2 1 1 [1013,1281,450,476] 1933-07 MACN Bernhauer & Chicago & Bernhauer Collection Bernhauer Bosq. Don. Bruch true Anfama 25 26 1 MLPA 25 26 5 1 2 [402,1084,487,514] 1973-08-24 IMLA L. Stange Tacanas Trancas 25 26 1 1 MLPA Bernhauer Bernhauer Richter 25 26 3 [398,918,562,588] 1908-01-27 MACN Bruch 25 26 2 [931,1178,562,588] 1908-01-27 MLPA Bruch true 25 26 1 1973-12 IMLA A. Sobral Dto. Rio Lipco Los Toldos 25 26 1 1 Salta