Maenioceras cf. crassum Bensaïd 1974: 105 Afromaenioceras Ammonoids of the Middle Devonian family Maenioceratidae in the Anti-Atlas of Morocco Bockwinkel, Jürgen Korn, Dieter European Journal of Taxonomy 2024 2024-01-30 921 1 1 35 Bockwinkel & Korn, 2024 Bockwinkel & Korn 2024 [586,903,1325,1352] Cephalopoda Maenioceratidae Afromaenioceras Animalia Agoniatitida 20 21 Mollusca species brumale sp. nov.   urn:lsid:zoobank.org:act: A55F968F-58F4-48D2-9D54-B3CD46403010  Fig. 11; Table 8   ?  Maeniocerascf. crassum–  Bensaïd 1974: 105, pl. 1 fig. 10, 10a. ?  Afromaenioceras n. sp.– Becker et al. 2004: pl. 1 figs 7–8.    Diagnosis Species of  Afromaenioceraswith thinly pachyconic and subevolute conch at 6 mmdm (ww/dm~ 0.70; uw/dm~ 0.38); conch thickly discoidal and subinvolute at 16 mmdm (ww/dm~0.55; uw/dm~0.18) with weakly depressed whorl profile (ww/wh ~ 1.15) and low coiling rate (WER ~1.55). Whorl profile at 16 mmconch diameter C-shaped with convex, weakly convergent flanks, broadly rounded ventrolateral shoulder and rounded venter. Ventrolateral shoulder with faint spiral groove. Growth lines coarse, strongly biconvex. Suture line with wide external lobe, small, subangular E 2lobe, inverted U-shaped ventrolateral saddle and symmetric, blunt V-shaped lateral lobe.    Etymology After the Latin adjective ‘ brumalis, - e’ = ‘summery’; the conch resembles a summer tire.    Type material    Holotype  MOROCCO• Anti-Atlas, Oufrane-E; bed 12, middle Givetian; Bockwinkeland Ebbighausen2002– 2004 Coll.; MB.C.31960(illustrated in Fig. 11A).    Paratypes  MOROCCO• 25specs; Anti-Atlas, Oued Mzerreb-W; beds 10/11, middle Givetian; Bockwinkeland Ebbighausen2004 Coll.; MB.C.31962.1 to MB.C.31962.25•  1 spec.; Anti-Atlas, Tafilalt, Jebel Ouaoufilal; middle Givetian; Bockwinkeland Ebbighausen2004 Coll.; MB.C.31961•  6specs; Anti-Atlas, Oufrane-E; bed 11, middle Givetian; Bockwinkeland Ebbighausen2004 Coll.; MB.C.31963.1 to MB.C.31963.6•  1 spec.; Anti-Atlas, Tiguisselt; bed 10, middle Givetian; Bockwinkeland Ebbighausen2004 Coll.; MB.C.31964.    Description  HolotypeMB.C.31960 is a very well-preserved limonitic internal mould with 16 mmconch diameter ( Fig. 11A). It is a thickly discoidal, subinvolute conch (ww/dm = 0.55; uw/dm = 0.18) with a weakly depressed whorl profile (ww/wh = 1.16) and a low coiling rate (WER = 1.53); flanks and venter are broadly rounded. The ventrolateral shoulder has a faint spiral groove. Traces of coarse growth lines are visible on the flanks. The last volution of the phragmocone has only ten chambers.  ParatypeMB.C.31961 is a thickly discoidal, subinvolute conch with 16 mmconch diameter and shows, like the two specimensdescribed above, rows of Housean Pits in the outer flank region ( Fig. 11B).   Fig. 11.  Afromaenioceras brumale sp. nov. A. Holotype MB.C.31960 from Oufrane-E, bed 12. B. Paratype MB.C.31961 from, Jebel Ouaoufilal. C. Paratype MB.C.31962.1 from Oued Mzerreb-W, beds 10/11. D. Paratype MB.C.31962.2 from Oued Mzerreb-W, bed 10/11; suture line at wh = 4.5 mm. E. Paratype MB.C.31962.3 from Oued Mzerreb-W, beds 10/11; suture line at wh = 3.8 mm. F. Paratype MB.C.31962.4 from Oued Mzerreb-W, bed 10/11; cross section. Scale bar units = 1 mm.   Table 8.Conch dimensions (in mm) and ratios of  Afromaenioceras brumale sp. nov.     specimen  dm  ww  wh  uw  ah  ww/dm  ww/wh  uw/dm  WER  IZR  MB.C.31960 16.2 9.0 7.7 2.9 3.1 0.55 1.16 0.18 1.53 0.60  MB.C.31961 15.8 8.9 7.7 3.1 – 0.57 1.16 0.20 – –  MB.C.31962.1 13.9 7.9 6.7 2.5 3.0 0.57 1.17 0.18 1.63 0.55  MB.C.31962.4 8.22 5.45 3.24 2.80 1.82 0.66 1.68 0.34 1.65 0.44  MB.C.31962.4 6.40 4.38 2.19 2.43 1.40 0.69 2.00 0.38 1.64 0.36  MB.C.31962.4 5.00 3.81 1.78 1.91 1.24 0.76 2.14 0.38 1.77 0.30  ParatypeMB.C.31962.4 was sectioned ( Fig. 11F) and shows two volutions up to a conch diameter of 8.2 mm. Between 5 and 8 mmconch diameter, the whorl profile changes from crescent-shaped to C-shaped and the conch becomes more slender and narrowly umbilicate.  ParatypesMB.C.31962.2 ( Fig. 11D) and MB.C.31962.3 ( Fig. 11E) Suture lines were drawn from those at wh = 4.5 and wh = 3.8 mm, respectively. Both show the Y-shaped external lobe with stair-shape flanks. However, they differ in the shape of the ventrolateral saddle, which is narrower in paratypeMB.C.31962.2, and in the shape of the lateral lobe, which is blunt V-shaped in paratypeMB.C.31962.2, but nearly U-shaped in paratypeMB.C.31962.3.    Remarks   Afromaenioceras crassumis a similar species but differs in the stouter conch (ww/dm~0.65; ww/wh~1.45; uw/dm 0.24) from  A. brumale sp. nov.(ww/dm~0.55; ww/wh ~1.15; uw/dm 0.25) at~ 15 mmdm. 4513871317 MB Bockwinkel & Ebbighausen Morocco Ebbighausen Anti-Atlas Bockwinkel 21 22 MB.C.31960 1 holotype 4513871302 MB Bockwinkel & Ebbighausen Morocco Anti-Atlas Givetian 21 22 MB.C.31962.1, MB.C.31962.2, MB.C.31962.3, MB.C.31962.4, MB.C.31962.5, MB.C.31962.6, MB.C.31962.7, MB.C.31962.8, MB.C.31962.9, MB.C.31962.10, MB.C.31962.11, MB.C.31962.12, MB.C.31962.13, MB.C.31962.14, MB.C.31962.15, MB.C.31962.16, MB.C.31962.17, MB.C.31962.18, MB.C.31962.19, MB.C.31962.20, MB.C.31962.21, MB.C.31962.22, MB.C.31962.23, MB.C.31962.24, MB.C.31962.25 25 paratype 4513871314 MB Bockwinkel & Ebbighausen Morocco Anti-Atlas Givetian 21 22 MB.C.31961 1 paratype 4513871332 MB Bockwinkel & Ebbighausen Morocco Anti-Atlas Givetian 21 22 MB.C.31963.1, MB.C.31963.2, MB.C.31963.3, MB.C.31963.4, MB.C.31963.5, MB.C.31963.6 6 paratype 4513871335 MB Bockwinkel & Ebbighausen Morocco Anti-Atlas Givetian 21 22 MB.C.31964 1 paratype