Laboulbenia anceps Peyr. ( Peyritsch 1873: 247 )
Laboulbenia elongata Thaxt. ( Thaxter 1890: 10 )
Laboulbenia lepida Thaxt. ( Thaxter 1896: 323 )
Laboulbenia anisodactyli Speg. ( Spegazzini 1914: 46 )
Laboulbenia pierantonii S.Colla. ( Colla 1926: 172 )
Laboulbenia rougetii var. europhili Siemaszko & W.Siemaszko ( Siemaszko & Siemaszko 1928: 203 )
Laboulbeniomycetes (Fungi, Ascomycota) of Denmark
Santamaria, Sergi
Pedersen, Jan
European Journal of Taxonomy
2021
2021-11-26
781
1
425
3RK8P
Peyr.
[624,964,1783,1810]
Laboulbeniomycetes
Laboulbeniaceae
Laboulbenia
Fungi
Laboulbeniales
152
153
Ascomycota
species
flagellata
MB#193832 Fig. 32
Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Wien. Mathematischnaturwissenschaftliche Classe. Abteilung 168: 247 ( Peyritsch 1873).–
Type: [Not designated] [Ind. loc.] [ Typelost?!] “Der Pilz befällt die Flügeldecken von Bembidium lunatumDuft., … Ausserdem habe ich ihn auf Anchomenus albipesF., and A. marginatusL. angetroffen”. [ Austria, according to Balazuc 1974b]
Laboulbenia ancepsPeyr. ( Peyritsch 1873: 247)[MB#172583] Laboulbenia elongataThaxt. ( Thaxter 1890: 10)[MB#187192] Laboulbenia lepidaThaxt. ( Thaxter 1896: 323)[MB#222474] Laboulbenia anisodactyliSpeg. ( Spegazzini 1914: 46)[MB#150969] Laboulbenia pierantoniiS.Colla. ( Colla 1926: 172)[Not registered in MycoBank] Laboulbenia rougetiivar. europhiliSiemaszko & W.Siemaszko ( Siemaszko & Siemaszko 1928: 203)[MB#139517]
Diagnostic features Cell V obtriangular, half of the length of cell IV, not connected with cell III. Outer appendage with the lower dorsal margin ± obscured; once branched above the suprabasal cell. Inner appendage consisting of a basal cell slightly shorter and narrower than basal cell of outer appendage, supporting a variable number of sterile and fertile branches bearing paired lateral antheridia. [Detailed descriptions: Sugiyama 1973; Majewski 1994b; Santamaria 1998; Villarreal et al.2010; De Kesel & Gerstmans 2011]
Distribution and hosts Laboulbenia flagellatais a very common and cosmopolitan species found on the five continents, infecting not less than 12 tribes and more than 80 genera of ground beetles ( Santamaria et al.1991; Majewski 2008; Haelewaters et al.2019b). Early reported in Denmarkby Rostrup (1916, 1935).
Collections examined from Denmark On Agonum emarginatum(Gyllenhal, 1827)(Col. Carabidae) DENMARK– Bornholm(B)• Raghammer Odde; 55°1.193′ N, 14°55.777′ E; VA99; 22 Jun. 2018; JP1151; JPdet.; ZMUC C-F-123675. On Agonum fuliginosum(Panzer, 1809)(Col. Carabidae) DENMARK– Bornholm(B)• Skelsmyrei Rønne Plantage; 55°4.375′ N, 14°45.096′ E; VB80; 25 Apr. 2014; JP507; JPdet.; ZMUC C-F-122990• Svartingedalenved Baggeå; 55°11.740′ N, 14°45.260′ E; VB81; 23 Jun. 2018; JP1161; JPdet.; ZMUC C-F-123687. – Østjylland(EJ)• Hald Ege; 56°24.287′ N, 9°20.550′ E; NH25; 19 Oct. 2013; JP794; JPdet.; ZMUC C-F-123289• Hampen Sø; 56°1.082′ N, 9°23.149′ E; NH20; 21 Mar. 2014; JP437; JPdet.; ZMUC C-F-122922• Kjellerup; 56°17.644′ N, 9°25.850′ E; NH23; 16 Feb. 2018; JP977; JPdet.; ZMUC C-F-123484• Ørnsøved Silkeborg; 56°9.446′ N, 9°31.488′ E; NH32; 11 Mar. 2017; JP277; JPdet.; ZMUC C-F-122756. – Fyn(F)• Gyldenstensinddaemmede Strand; 55°34.447′ N, 10°7.211′ E; NG75; 2Apr. 2014; JP707; JPdet.; ZMUC C-F-123197• Rishave; 55°33.475′ N, 10°24.013′ E; NG85; 19 Apr. 2018; JP1071; JPdet.; ZMUC C-F-123588. – Lolland, Falster, Møn(LFM)• Lysemoseved Maribo; 54°46.971′ N, 11°28.959′ E; PF57; 7 May 2017; JP660; JPdet.; ZMUC C-F-123149• Nyord; 55°2.885′ N, 12°12.380′ E; UB20; 27 Apr. 2013; JP830; JPdet.; ZMUC C-F-123324• Vestfor Bandholm; 54°50.303′ N, 11°28.050′ E; PF57; 3 Jan. 2019; JP1306; JPdet.; ZMUC C-F-123841. – Nordøstjylland(NEJ)• Vesterskoveni Tofte Skov; 56°50.346′ N, 10°13.361′ E; NJ70; 7 Aug. 2013; JP316; JPdet.; ZMUC C-F-122795. – Nordøstsjaelland(NEZ)• Eriksholm Skov; 55°41.771′ N, 11°47.912′ E; PG77; 4 Nov. 2017; JP735; JPdet.; ZMUC C-F-123228• Faergelunden; 55°50.540′ N, 12°1.735′ E; UB19; 16 Nov. 2013; JP594; JPdet.; ZMUC C-F-123083• Gurre Vang; 56°1.470′ N, 12°29.486′ E; UC41; 14 Apr. 2017; JP262; JPdet.; ZMUC C-F-122741• Indelukketved Frederiksborg Slot; 55°56.191′ N, 12°17.861′ E; UC30; 19 Mar. 2017; JP664; JPdet.; ZMUC C-F-123153• ibid.; 12 Apr. 2016; JP18; JPdet.; ZMUC C-F-122491• Nordfor Risø; 55°42.184′ N, 12°6.692′ E; UB17; 14 Dec. 2013; R. Aagaard Jensen631; JPdet.; ZMUC C-F-123120• Vaserne; 55°49.129′ N, 12°26.343′ E; UB38; 21 Apr. 2018; L. Thomas1119; JPdet.; ZMUC C-F-123639. – Nordvestjylland(NWJ)• Vestfor Vinderup; 56°28.727′ N, 8°44.810′ E; MH85; 19 Jan. 2015; JP773; JPdet.; ZMUC C-F-123268. – Nordvestsjaelland(NWZ)• Bognaes Skovpå Tuse Naes; 55°44.966′ N, 11°45.817′ E; PG78; 10 Dec. 2013; JP881; JPdet.; ZMUC C-F-123379• Vesterlyng; 55°44.195′ N, 11°17.276′ E; PG47; 9 Feb. 2014; JP642; JPdet.; ZMUC C-F-123130. – Sydjylland(SJ)• Lakolk; 55°8.325′ N, 8°29.618′ E; MG61; 31 May 2013; JP291; JPdet.; ZMUC C-F-122770. – Sydsjaelland(SZ)• Denderup Søi Denderup Vaenge; 55°15.075′ N, 11°57.366′ E; PG82; 5 May 2016; JP489; JPdet.; ZMUC C-F-122973• Gammel Kalvehave; 54°59.745′ N, 12°8.232′ E; UA19; 25 Apr. 2019; JP1443; JPdet.; ZMUC C-F-123991• Østerskovved Langebaek; 54°59.235′ N, 12°6.280′ E; UA19; 14 Jan. 2012; JP544; JPdet.; ZMUC C-F-123028• Praestø; 55°6.913′ N, 12°2.035′ E; UB11; 25 Apr. 2019; JP1437; JPdet.; ZMUC C-F-123984. – Vestjylland(WJ)• Velling; 56°3.078′ N, 8°18.596′ E; MH51; 16 Dec. 2018; JP1254; JPdet.; ZMUC C-F-123782. Fig. 32. A–G. Laboulbenia flagellataPeyr. Maturethalli from Agonum fuliginosum(A), Agonum muelleri(B), Loricera pilicornis(C), Anisodactylus binotatus(D), Platynus assimilis(E–F), and Anchomenus dorsalis(G). Scale bars: 50 µm. Photographs from slides ZMUC C-F-122491 (A), ZMUC C-F-122590 (B), ZMUC C-F-123321 (C), ZMUC C-F-122589 (D), ZMUC C-F-122578 (E–F), ZMUC C-F-122829 (G). On Agonum gracileSturm, 1824(Col. Carabidae) DENMARK– Nordøstjylland(NEJ)• Tranesigi Tofte Skov; 56°51.597′ N, 10°13.575′ E; NJ70; 1 Oct. 2010; JP74; JPdet.; ZMUC C-F-122549. On Agonum marginatum(Linnaeus, 1758)(Col. Carabidae) DENMARK– Nordøstsjaelland(NEZ)• Nordhavnstippenpå Københavns Nordhavn; 55°43.091′ N, 12°37.088′ E; UB57; 29 Apr. 2018; JP1090; JPdet.; ZMUC C-F-123608• Udfor Kongelundsfortet; 55°33.724′ N, 12°33.731′ E; UB46; 15 Apr. 2018; JP1051; JPdet.; ZMUC C-F-123567. – Sydjylland(SJ)• Sønderstrandpå Rømø; 55°6.373′ N, 8°29.380′ E; MG60; 25 Oct. 2017; H. Liljehult619; JPdet.; ZMUC C-F-123107. On Agonum micans(Nicolai, 1822)(Col. Carabidae) DENMARK– Fyn(F)• Kratholmved Bellinge; 55°19.710′ N, 10°19.335′ E; NG83; 18 May 2016; JP87; JPdet.; ZMUC C-F-122562. On Agonum muelleri(Herbst, 1784)(Col. Carabidae) DENMARK– Østjylland(EJ)• Kjellerup; 56°17.644′ N, 9°25.850′ E; NH23; 16 Feb. 2018; JP978; JPdet.; ZMUC C-F-123485. – Nordøstsjaelland(NEZ)• Brønshøj; 55°42.234′ N, 12°30.135′ E; UB47; 22 Jun. 2016; JP113; JPdet.; ZMUC C-F-122590• Naerum; 55°49.077′ N, 12°32.686′ E; UB48; 2 Dec. 2011; JP474; JPdet.; ZMUC C-F-122958. – Nordvestjylland(NWJ)• Legind; 56°45.832′ N, 8°48.782′ E; MH89; 19 Oct. 2008; JP75; JPdet.; ZMUC C-F-122550. – Nordvestsjaelland(NWZ)• Nordbreddenaf Skarresø; 55°39.469′ N, 11°23.147′ E; PG47; 18 Feb. 2017; JP340; JPdet.; ZMUC C-F-122819. On Agonum piceum(Linnaeus, 1758)(Col. Carabidae) DENMARK– Nordøstjylland(NEJ)• Sydfor Krogsagergård; 56°37.534′ N, 9°45.551′ E; NH47; 24 Nov. 2019; JP1556; JPdet.; ZMUC C-F-124317. – Vestjylland(WJ)• Velling; 56°3.078′ N, 8°18.596′ E; MH51; 16 Dec. 2018; JP1248; JPdet.; ZMUC C-F-123776. On Agonum thoreyiDejean, 1828(Col. Carabidae) DENMARK– Bornholm(B)• Svartingedalenved Baggeå; 55°11.740′ N, 14°45.260′ E; VB81; 23 Jun. 2018; JP1160; JPdet.; ZMUC C-F-123686. – Østjylland(EJ)• Viemoseved Ringkloster; 56°0.632′ N, 9°57.400′ E; NH50; 12 Mar. 2017; JP531; JPdet.; ZMUC C-F-123015. – Fyn(F)• Gyldenstensinddaemmede Strand; 55°34.447′ N, 10°7.211′ E; NG75; 2 Apr. 2014; JP708; JPdet.; ZMUC C-F-123198. – Lolland, Falster, Møn(LFM)• Nyord; 55°2.885′ N, 12°12.380′ E; UB20; 27 Apr. 2013; JP831; JPdet.; ZMUC C-F-123325. – Nordøstsjaelland(NEZ)• Auderød Skov; 55°59.117′ N, 12°5.579′ E; UC10; 28 Jan. 2018; J. Søgaard Hansen997; J. Søgaard Hansendet.; ZMUC C-F-123504• Brønshøj( Degnemosen); 55°42.043′ N, 12°30.352′ E; UB47; 26 May 2016; JP289; JPdet.; ZMUC C-F-122768• Faergelunden; 55°50.540′ N, 12°1.735′ E; UB19; 16 Nov. 2013; JP595; JPdet.; ZMUC C-F-123084• Lyngenved Ølstykke; 55°47.738′ N, 12°10.253′ E; UB28; 15 May 2013; JP862; JPdet.; ZMUC C-F-123357• Udfor Kongelundsfortet; 55°33.724′ N, 12°33.731′ E; UB46; 15 Apr. 2018; JP1052; JPdet.; ZMUC C-F-123568. – Nordvestjylland(NWJ)• Nordfor Landtingved Vinderup; 56°30.049′ N, 8°45.384′ E; MH86; 19 Jan. 2015; JP566; JPdet.; ZMUC C-F-123052. – Sydjylland(SJ)• Lakolk; 55°8.325′ N, 8°29.618′ E; MG61; 31 May 2013; JP292; JPdet.; ZMUC C-F-122771. – Sydsjaelland(SZ)• Even Bro; 55°8.694′ N, 12°0.601′ E; UB11; 17 Feb. 2019; JP1365; JPdet.; ZMUC C-F-123905• Gammel Kalvehave; 54°59.745′ N, 12°8.232′ E; UA19; 25 Apr. 2019; JP1442; JPdet.; ZMUC C-F-123990• Østerskovved Langebaek; 54°59.235′ N, 12°6.280′ E; UA19; 14 Jan. 2012; JP545; JPdet.; ZMUC C-F-123029• ibid.; 54°59.572′ N, 12°6.715′ E; UA19; 3 Jan. 2019; JP1273; JPdet.; ZMUC C-F-123803. On Agonum versutumSturm, 1824(Col. Carabidae) DENMARK– Bornholm(B)• Skelsmyrei Rønne Plantage; 55°4.375′ N, 14°45.096′ E; VB80; 25 Apr. 2014; JP505; JPdet.; ZMUC C-F-122988. On Agonum viduum(Panzer, 1797)(Col. Carabidae) DENMARK– Lolland, Falster, Møn(LFM)• Lysemoseved Maribo; 54°46.971′ N, 11°28.959′ E; PF57; 7 May 2017; JP659; JPdet.; ZMUC C-F-123148. On Anchomenus dorsalis(Pontoppidan, 1763)(Col. Carabidae) DENMARK– Bornholm(B)• Østfor Østerlars; 55°9.619′ N, 14°59.763′ E; VB91; 25 Apr. 2014; JP210; JPdet.; ZMUC C-F-122689. – Nordøstsjaelland(NEZ)• Brøndby; 55°39.089′ N, 12°24.487′ E; UB37; 22 Apr. 2016; JP45; JPdet.; ZMUC C-F-122520. – Sydsjaelland(SZ)• Holtug Kalkbrud; 55°20.470′ N, 12°26.678′ E; UB33; 21 Sep. 2013; JP347; JPdet.; ZMUC C-F-122829. On Anisodactylus binotatus(Fabricius, 1787)(Col. Carabidae) DENMARK– Nordøstsjaelland(NEZ)• Utterslev Mose; 55°42.714′ N, 12°29.907′ E; UB47; 3 Jun. 2016; H.S. Pedersen112; JPdet.; ZMUC C-F-122589. On Harpalus affinis(Schrank, 1781)(Col. Carabidae) DENMARK– Østjylland(EJ)• Kjellerup; 56°17.282′ N, 9°26.197′ E; NH23; 16 Oct. 2014; JP320; JPdet.; ZMUC C-F-122798. – Nordøstsjaelland(NEZ)• Naerum; 55°49.077′ N, 12°32.686′ E; UB48; 2 Dec. 2011; JP476; JPdet.; ZMUC C-F-122960. On Loricera pilicornis(Fabricius, 1775)(Col. Carabidae) DENMARK– Lolland, Falster, Møn(LFM)• Nyord; 55°2.885′ N, 12°12.380′ E; UB20; 27 Apr. 2013; JP827; JPdet.; ZMUC C-F-123321. – Nordøstsjaelland(NEZ)• Brønshøj( Degnemosen); 55°42.043′ N, 12°30.352′ E; UB47; 26 May 2016; JP288; JPdet.; ZMUC C-F-122767• Gurre Vang; 56°1.470′ N, 12°29.486′ E; UC41; 14 Apr. 2017; JP265; JPdet.; ZMUC C-F-122744. – Sydsjaelland(SZ)• Flommenved Sorø; 55°25.957′ N, 11°34.477′ E; PG64; 25 Jul. 2009; JP77; JPdet.; ZMUC C-F-122552• Sorø, Kristiansminde; 55°25.115′ N, 11°35.209′ E; PG64; 22 Jul. 2013; H. Enghoff308; JPdet.; ZMUC C-F-122787. On Nebria brevicollis(Fabricius, 1792)(Col. Carabidae) DENMARK– Nordøstsjaelland(NEZ)• Esrum Lund; 56°2.938′ N, 12°22.141′ E; UC31; 1–12 May 2006; J. Høier1571; P. Jørum; ZMUC C-F-124332• Lille Hareskovved Harrevads Bro; 55°45.966′ N, 12°22.601′ E; UB38; 1–12 May 2006; J. Høier1573; P. Jørum; ZMUC C-F-124334• Nørreskoven; 55°47.865′ N, 12°23.054′ E; UB38; 1–12 Sep. 2006; J. Høier1572; P. Jørum; ZMUC C-F-124333. On Oxypselaphus obscurus(Herbst, 1784)(Col. Carabidae) DENMARK– Østjylland(EJ)• Hampen Sø; 56°1.082′ N, 9°23.149′ E; NH20; 21 Mar. 2014; JP433; JPdet.; ZMUC C-F-122918. – Fyn(F)• Rishave; 55°33.475′ N, 10°24.013′ E; NG85; 19 Apr. 2018; JP1070; JPdet.; ZMUC C-F-123587. – Lolland, Falster, Møn(LFM)• Krenkerup Haveskov; 54°46.408′ N, 11°39.955′ E; PF77; 8 Jul. 1993; S. Langemark& O.E. Meyer907; JPdet.; ZMUC C-F-123405• ibid.; 19 Jul. 1993; S. Langemark& O.E. Meyer922; JPdet.; ZMUC C-F-12342• ibid.; 16–24 Aug. 1993; S. Langemark& O.E. Meyer912; JPdet.; ZMUC C-F-123410• Lysemoseved Maribo; 54°46.971′ N, 11°28.959′ E; PF57; 7 May 2017; JP653; JPdet.; ZMUC C-F-123142• ibid.; 4 Jun. 2018; JP1126; JPdet.; ZMUC C-F-123648• Nyord; 55°2.885′ N, 12°12.380′ E; UB20; 27 Apr. 2013; JP829; JPdet.; ZMUC C-F-123323• Ulvshale; 55°2.636′ N, 12°15.638′ E; UB20; 26 May 2018; JP1109; JPdet.; ZMUC C-F-123627• Vestfor Bandholm; 54°50.303′ N, 11°28.050′ E; PF57; 3 Jan. 2019; JP1309; JPdet.; ZMUC C-F-123844. – Nordøstsjaelland(NEZ)• Gurre Vang; 56°1.470′ N, 12°29.486′ E; UC41; 14 Apr. 2017; H. Enghoff267; JPdet.; ZMUC C-F-122746• Haettingsholm; 55°49.260′ N, 12°13.967′ E; UB38; 21 Apr. 2018; L. Thomas1118; JPdet.; ZMUC C-F-123638• Nordfor Frederikssund; 55°51.404′ N, 12°3.534′ E; UB17; 14 Dec. 2013; R. Aagaard Jensen407; JPdet.; ZMUC C-F-122891• Stampeskovved Rådvad; 55°48.332′ N, 12°33.138′ E; UB48; 14 Apr. 2013; JP780; JPdet.; ZMUC C-F-123275• Vestfor Trørød; 55°50.223′ N, 12°31.185′ E; UB49; 30 Mar. 2019; JP1404; JPdet.; ZMUC C-F-123950. – Nordvestsjaelland(NWZ)• Bognaes Skovpå Tuse Naes; 55°44.966′ N, 11°45.817′ E; PG78; 10 Dec. 2013; JP880; JPdet.; ZMUC C-F-123378• Nordbreddenaf Skarresø; 55°39.469′ N, 11°23.147′ E; PG47; 18 Feb. 2017; JP339; JPdet.; ZMUC C-F-122818. – Sydjylland(SJ)• Draved Skov; 55°0.964′ N, 8°58.395′ E; MF99; 1 Jun. 2013; JP304; JPdet.; ZMUC C-F-122783• Østerskovved Langebaek; 54°59.235′ N, 12°6.280′ E; UA19; 14 Jan. 2012; JP539; JPdet.; ZMUC C-F-123023• Sølsted Mose; 55°1.829′ N, 8°50.351′ E; MF89; 2 Jun. 2013; JP488; JPdet.; ZMUC C-F-122972. – Sydsjaelland(SZ)• Gammel Kalvehave; 54°59.745′ N, 12°8.232′ E; UA19; 24 Mar. 2018; JP1004; JPdet.; ZMUC C-F-123511• ibid.; 27 Apr. 2018; JP1082; JPdet.; ZMUC C-F-123600• Østerskovved Langebaek; 54°59.572′ N, 12°6.715′ E; UA19; 3 Jan. 2019; JP1278; JPdet.; ZMUC C-F-123808• Storskovsyd for Sparresholm; 55°13.234′ N, 11°57.463′ E; PG82; 17 Feb. 2019; JP1386; JPdet.; ZMUC C-F-123929. On Paranchus albipes(Fabricius, 1796)(Col. Carabidae) DENMARK– Bornholm(B)• Døndalensudløb; 55°13.656′ N, 14 53.078′ E; VB92; 24 Jun. 2018; JP1143; JPdet.; ZMUC C-F-123666. – Nordøstsjaelland(NEZ)• Kongensbøge, Gribskov; 56°0.755′ N, 12°22.033′ E; UC13; 20 Apr. 2018; L. Kjaer-Thomsen1116; JPdet.; ZMUC C-F-123636. On Platynus assimilis(Paykull, 1790)(Col. Carabidae) DENMARK– Bornholm(B)• Døndal; 55°13.489′ N, 14 52.936′ E; VB92; 26 May 2017; L.K. Thomsen& L. Thomas426; JPdet.; ZMUC C-F-122911• Svartingedalenved Baggeå; 55°11.740′ N, 14°45.260′ E; VB81; 23 Jun. 2018; JP1157; JPdet.; ZMUC C-F-123682. – Lolland, Falster, Møn(LFM)• Keldskov; 54°40.114′ N, 11°33.363′ E; PF66; 15 Jun. 2012; H. Liljehult44; JPdet.; ZMUC C-F-122519• Krenkerup Haveskov; 54°46.408′ N, 11°39.955′ E; PF77; 5 May 1993; S. Langemark& O.E. Meyer895; JPdet.; ZMUC C-F-123393• ibid.; 12 May 1993; S. Langemark& O.E. Meyer104; JPdet.; ZMUC C-F-122579• ibid.; 25 May 1993; S. Langemark& O.E. Meyer909; JPdet.; ZMUC C-F-123407• ibid.; 16 Jun. 1993; S. Langemark& O.E. Meyer103; JPdet.; ZMUC C-F-122578• Ulvshale; 55°2.636′ N, 12°15.638′ E; UB20; 26 May 2018; JP1108; JPdet.; ZMUC C-F-123626. – Nordøstjylland(NEJ)• Engskoveni Tofte Skov; 56°51.525′ N, 10°15.065′ E; NJ70; 8–22 Aug. 2013; I. Aggerholm935; JPdet.; ZMUC C-F-123435. – Nordøstsjaelland(NEZ)• Herthadal, Lejre; 55°36.853′ N, 11°56.621′ E; PG86; 20 May 2018; L. Kjaer-Thsen1117; L. Kjaer-Thomsendet.; ZMUC C-F-123637• Lille Lyngby Mose; 55°40.831′ N, 12°29.094′ E; UC20; 28 Jan. 2018; J. Søgaard Hansen998; J. Søgaard Hansendet.; ZMUC C-F-123505. – Nordvestsjaelland(NWZ)• Nordøstbreddenaf Tissø; 55°35.612′ N, 11°18.461′ E; PG46; 1 May 2013; JP757; JPdet.; ZMUC C-F-123251• Nordbreddenaf Skarresø; 55°39.425′ N, 11°22.998′ E; PG47; 1 May 2013; JP855; JPdet.; ZMUC C-F-123350. – Sydsjaelland(SZ)• Åshøje Overdrev; 55°26.368′ N, 12°6.359′ E; UB14; 6 Oct. 2018; L. Kjaer-Thomsen& L. Thomas1235; JPdet.; ZMUC C-F-123762• Østerskovved Langebaek; 54°59.331′ N, 12°6.357′ E; UA19; 27 Mar. 2018; JP1020; JPdet.; ZMUC C-F-123527• ibid.; 3 Jan. 2019; JP1277; JPdet.; ZMUC C-F-123807• Sorø, Kristiansminde; 55°25.115′ N, 11°35.209′ E; PG64; 22 Jul. 2013; H. Enghoff307; JPdet.; ZMUC C-F-122786• ibid.; 2 Jul. 2014; JP382; JPdet.; ZMUC C-F-122865. On Platynus livens(Gyllenhal, 1810)(Col. Carabidae) DENMARK– Bornholm(B)• Skelsmyrei Rønne Plantage; 55°4.375′ N, 14°45.096′ E; VB80; 25 Apr. 2014; H. Liljehult231; JPdet.; ZMUC C-F-122712. – Sydsjaelland(SZ)• Holmegårds Mose; 55°17.685′ N, 11°47.928′ E; PG73; 31 Mar. 2019; JP1418; JPdet.; ZMUC C-F-123964. On Pterostichus oblongopunctatus(Fabricius, 1787)(Col. Carabidae) DENMARK– Nordøstsjaelland(NEZ)• Jonstrup Vangved Store Sejben Mose; 55°45.452′ N, 12°22.217′ E; UB38; 1–12 Jul. 2006; J. Høier1575; P. Jørumdet.; ZMUC C-F-124336• Lille Hareskovved Harrevads Bro; 55°45.966′ N, 12°22.601′ E; UB38; 1–12 May 2006; J. Høier1574; P. Jørumdet.; ZMUC C-F-124335. On Pterostichus vernalis(Panzer, 1796)(Col. Carabidae) DENMARK– Bornholm(B)• Østfor Østerlars; 55°9.619′ N, 14°59.763′ E; VB91; 25 Apr. 2014; JP211; JPdet.; ZMUC C-F-122690. – Lolland, Falster, Møn(LFM)• Vestfor Bandholm; 54°50.303′ N, 11°28.050′ E; PF57; 3 Jan. 2019; JP1326; JPdet.; ZMUC C-F-123865. – Nordøstsjaelland(NEZ)• Utterslev Mose; 55°42.714′ N, 12°29.907′ E; UB47; 28 Oct. 2012; JP43; JPdet.; ZMUC C-F-122518. – Nordvestsjaelland(NWZ)• Bognaes Skovpå Tuse Naes; 55°44.966′ N, 11°45.817′ E; PG78; 10 Dec. 2013; JP877; JPdet.; ZMUC C-F-123374• Flaskennord for Reersø; 55°31.965′ N, 11°9.831′ E; PG35; 26 Jan. 2014; JP612; JPdet.; ZMUC C-F-123100• Flyndersø ved Dybesø; 55°58.151′ N, 11°45.621′ E; PH60; 4 Nov. 2017; JP718; JPdet.; ZMUC C-F-123210• Nordbreddenaf Skarresø; 55°39.469′ N, 11°23.147′ E; PG47; 18 Feb. 2017; JP338; JPdet.; ZMUC C-F-122817. – Sydsjaelland(SZ)• Gammel Kalvehave; 54°59.745′ N, 12°8.232′ E; UA19; 24 Mar. 2018; JP1001; JPdet.; ZMUC C-F-123508• ibid.; 27 Apr. 2018; JP1079; JPdet.; ZMUC C-F-123597• ibid.; 25 Apr. 2019; JP1444; JPdet.; ZMUC C-F-123992• Østerskovved Langebaek; 54°59.572′ N, 12°6.715′ E; UA19; 3 Jan. 2019; JP1290; JPdet.; ZMUC C-F-123822• Praestø; 55°6.913′ N, 12°2.035′ E; UB11; 25 Apr. 2019; JP1435; JPdet.; ZMUC C-F-123982.
Remarks Laboulbenia flagellatais a binomen associated with an awkward species, described with very loose characters, waiting for a typedesignation and a precise delimitation of characteristics. Translation from German Peyritsch’s (1873)protologue states: “bright yellowish brown, only with a blackened perithecial apex and base as well, appendages few in number (4–7), rather similar, unbranched or divided from the base, not tinged, typically exceeding the perithecial apex”. In short, characteristics shared by numerous species. Moreover, Peyritsch (1873)mentioned three hosts: Bembidium lunatum(sic) (a host which now typically has been mentioned yielding Laboulbenia vulgarisor Laboulbenia pedicellataor perhaps some related species but never L. flagellata), Anchomenus albipes(now Paranchus albipes) (which is now the typical host for L. collae, although mixed infections with L. flagellataare common according to De Kesel, pers. comm.), and lastly, Anchomenus marginatus(now Agonum marginatum). The genus Agonumseems to be acceptable as the only one of these three genera where we find what most authors have assumed over the years to be L. flagellata. Therefore, an epitypification is highly desirable, where this host or any related might be considered as the mostly appropriate. It is generally accepted by students of Laboulbenialesthat L. flagellatais a species complex. Over the years, laboulbeniologists have been delimiting the taxon only a little better, resulting in what we have today. However, the amount of synonyms (from five to nine, depending on the authors′ criteria) and around ten infraspecific taxa, lacking any relevance, have meanwhile proliferated to increase the confusion. De Kesel & VanDen Neucker (2005)studied the morphological plasticity of L. flagellataobtained from Agonumspp.and Loricera pilicornisrelative to environmental factors, concluding that there is no morphological evidence for segregation of their studied samples into different species. A very initial study with molecular techniques and morphometrics, using thalli from different species of ground beetles, demonstrated the existence of two clades ( Haelewaters et al.2019b). These two clades separate significantly thalli from Limodromus assimilis(Paykull, 1790)(= Platynus assimilis) from those on Agonum micansand Loricera pilicornis. Unfortunately the absence of typification for the species and the broadness of its concept reduce the impact of this study because it cannot be assured that what is sequenced is certainly a L. flagellatabecause of the absence of a fair morphological characterization including iconography in the protologue. We have had at our disposal very abundant material from Denmarkwhich has permitted us to perceive the enormous morphological variation of thalli within the concept of L. flagellata. Without being able to attain significant conclusions, we present some images which eloquently exemplify the status of the “species complex”. In Fig. 32photographs of fungi from Agonum fuliginosum( Fig. 32A), A. muelleri( Fig. 32B), Anchomenus dorsalis( Fig. 32G), Anisodactylus binotatus( Fig. 32D), Loricera pilicornis( Fig. 32C) and Platynus assimilis( Fig. 32E–F) have been included. Thalli from Agonumspp.fit what may be broadly accepted as a “typical” L. flagellata; in the middle we have the thalli from Anisodactylusand LoriceraLatreille, 1802showing some interesting differences; and finally we have thalli from Platynusand Anchomenusthat may represent future new species. Laboulbenia giganteaIstvánffi, which has been considered a synonym of L. flagellatafor a very long time, was reinstated in a recent study by Rossi & Christian (2020)
4988641704
Austria
153
154
Austria
152
153
1
type
4988642036
On Agonum emarginatum (Gyllenhal, 1827)
2018-06-22
ZMUC
JP
Denmark
JP
55.019882
Raghammer Odde
1
14.929617
153
154
ZMUC C-F-123675
1
Bornholm
4988642275
On Agonum fuliginosum (Panzer, 1809)
2014-04-25
ZMUC
JP
Denmark
JP
55.072918
Ronne Plantage
1
14.7516
Skelsmyre
153
154
ZMUC C-F-122990
1
Bornholm
4988640870
2018-06-23
ZMUC
JP
Denmark
JP
55.195667
Baggea
1
14.7543335
Svartingedalen
153
154
ZMUC C-F-123687
1
Bornholm
4988642025
2013-10-19
ZMUC
JP
Denmark
JP
56.404785
Hald Ege
1
9.3425
153
154
ZMUC C-F-123289
1
Ostjylland
4988642168
2014-03-21
ZMUC
JP
Denmark
JP
56.018032
Hampen So
1
9.385817
153
154
ZMUC C-F-122922
1
Ostjylland
4988640529
2018-02-16
ZMUC
JP
Denmark
JP
56.294067
Kjellerup
1
9.430834
153
154
ZMUC C-F-123484
1
Ostjylland
4988641548
2017-03-11
ZMUC
JP
Denmark
JP
56.157433
Silkeborg
1
9.5248
Ornso
153
154
ZMUC C-F-122756
1
Ostjylland
4988641010
2014-04-02
ZMUC
JP
Denmark
JP
55.574116
Strand
1
10.120183
Gyldenstens
153
154
ZMUC C-F-123197
1
Fyn
4988642338
2018-04-19
ZMUC
JP
Denmark
JP
55.557915
Rishave
1
10.400217
153
154
ZMUC C-F- 123588
1
Fyn
4988641409
2017-05-07
ZMUC
JP
Denmark
JP
54.78285
Maribo
1
11.48265
Lysemose
153
154
ZMUC C-F-123149
1
Lolland, Falster, Mon
4988641646
2013-04-27
ZMUC
JP
Denmark
JP
55.048084
Nyord
1
12.206333
153
154
ZMUC C-F-123324
1
Lolland, Falster, Mon
4988641757
2019-01-03
ZMUC
JP
Denmark
JP
54.838383
Bandholm
1
11.4675
Vest
153
154
ZMUC C-F-123841
1
Lolland, Falster, Mon
4988640416
2013-08-07
NEJ
JP
Denmark
JP
56.8391
Tofte Skov
1
10.222683
Vesterskoven
153
154
ZMUC C-F-122795
1
Nordostjylland
4988641821
2017-11-04
ZMUC
JP
Denmark
JP
55.696182
Eriksholm Skov
1
11.798533
153
154
ZMUC C-F-123228
1
Nordostsjaelland
4988640367
2013-11-16
ZMUC
JP
Denmark
JP
55.842335
Faergelunden
1
12.028916
153
154
ZMUC C-F-123083
1
Nordostsjaelland
4988641855
2017-04-14
ZMUC
JP
Denmark
JP
56.0245
Gurre Vang
1
12.491433
153
154
ZMUC C-F-122741
1
Nordostsjaelland
4988641889
2017-03-19
ZMUC
JP
Denmark
JP
55.936516
Frederiksborg Slot
1
12.297684
Indelukket
153
154
ZMUC C-F-123153
1
Nordostsjaelland
4988640994
[609,1268,1962,1989]
2016-04-12
ZMUC
JP
Denmark
JP
55.936516
Frederiksborg Slot
1
12.297684
Indelukket
153
154
ZMUC C-F-122491
1
Nordostsjaelland
4988640951
2013-12-14
ZMUC
R. Aagaard Jensen
Denmark
JP
155
156
55.703068
Riso
1
12.111533
Nord
153
154
ZMUC C-F- 123120
1
Nordostsjaelland
4988640898
2018-04-21
ZMUC
L. Thomas
Denmark
JP
55.818817
Vaserne
1
12.43905
155
156
ZMUC C-F-123639
1
Nordostsjaelland
4988640316
2015-01-19
ZMUC
JP
Denmark
JP
56.478783
Vinderup
1
8.746834
Vest
155
156
ZMUC C-F-123268
1
Nordvestjylland
4988640550
2013-12-10
ZMUC
JP
Denmark
JP
55.74943
Tuse Naes
1
11.763617
Bognaes Skov
155
156
ZMUC C-F-123379
1
Nordvestsjaelland
4988641256
2014-02-09
ZMUC
JP
Denmark
JP
55.736584
Vesterlyng
1
11.287933
155
156
ZMUC C-F-123130
1
Nordvestsjaelland
4988640667
2013-05-31
ZMUC
JP
Denmark
JP
55.13875
Lakolk
1
8.493633
155
156
ZMUC C-F-122770
1
Sydjylland
4988641123
2016-05-05
ZMUC
JP
Denmark
JP
55.25125
Denderup Vaenge
1
11.9561
Denderup So
155
156
ZMUC C-F-122973
1
Sydsjaelland
4988640848
2019-04-25
ZMUC
JP
Denmark
JP
54.99575
Gammel Kalvehave
1
12.1372
155
156
ZMUC C-F-123991
1
Sydsjaelland
4988641205
2012-01-14
ZMUC
JP
Denmark
JP
54.98725
Langebaek
1
12.104667
Osterskov
155
156
ZMUC C-F-123028
1
Sydsjaelland
4988641042
2019-04-25
ZMUC
JP
Denmark
JP
55.115215
Praesto
1
12.033916
155
156
ZMUC C-F-123984
1
Sydsjaelland
4988641606
2018-12-16
ZMUC
JP
Denmark
JP
56.0513
Velling
1
8.309934
155
156
ZMUC C-F-123782
1
Vestjylland
4988640950
On Agonum gracile Sturm, 1824
2010-10-01
NEJ
JP
Denmark
JP
56.85995
Tofte Skov
1
10.22625
Tranesig
155
156
ZMUC C-F-122549
1
Nordostjylland
4988641671
On Agonum marginatum (Linnaeus, 1758)
2018-04-29
ZMUC
JP
Denmark
JP
55.71818
Kobenhavns Nordhavn
1
12.618134
Nordhavnstippen
155
156
ZMUC C-F-123608
1
Nordostsjaelland
4988640493
2018-04-15
ZMUC
JP
Denmark
JP
55.562065
Kongelundsfortet
1
12.562183
Ud
155
156
ZMUC C-F-123567
1
Nordostsjaelland
4988641079
2017-10-25
ZMUC
H. Liljehult
Denmark
JP
55.106216
Romo
1
8.489667
Sonderstrand
155
156
ZMUC C-F-123107
1
Sydjylland
4988642255
On Agonum micans (Nicolai, 1822)
2016-05-18
ZMUC
JP
Denmark
JP
55.3285
Bellinge
1
10.32225
Kratholm
155
156
ZMUC C-F-122562
1
Fyn
4988642019
On Agonum muelleri (Herbst, 1784)
2018-02-16
ZMUC
JP
Denmark
JP
56.294067
Kjellerup
1
9.430834
155
156
ZMUC C-F-123485
1
Ostjylland
4988641730
2016-06-22
ZMUC
JP
Denmark
JP
55.7039
Bronshoj
1
12.50225
155
156
ZMUC C-F-122590
1
Nordostsjaelland
4988642227
2011-12-02
ZMUC
JP
Denmark
JP
55.81795
Naerum
1
12.544766
155
156
ZMUC C-F-122958
1
Nordostsjaelland
4988641542
2008-10-19
ZMUC
JP
Denmark
JP
56.763866
Legind
1
8.813033
155
156
ZMUC C-F-122550
1
Nordvestjylland
4988640790
2017-02-18
ZMUC
JP
Denmark
JP
55.657818
Skarreso
1
11.385783
Nordbredden
155
156
ZMUC C-F-122819
1
Nordvestsjaelland
4988640700
On Agonum piceum (Linnaeus, 1758)
2019-11-24
NEJ
JP
Denmark
JP
56.62557
Krogsagergard
1
9.759183
Syd
155
156
ZMUC C-F-124317
1
Nordostjylland
4988640784
2018-12-16
ZMUC
JP
Denmark
JP
56.0513
Velling
1
8.309934
155
156
ZMUC C-F-123776
1
Vestjylland
4988640618
On Agonum thoreyi Dejean, 1828
2018-06-23
ZMUC
JP
Denmark
JP
55.195667
Baggea
1
14.7543335
Svartingedalen
155
156
ZMUC C-F-123686
1
Bornholm
4988640518
2017-03-12
ZMUC
JP
Denmark
JP
56.010532
Ringkloster
1
9.956667
Viemose
155
156
ZMUC C-F-123015
1
Ostjylland
4988641872
2014-04-02
ZMUC
JP
Denmark
JP
55.574116
Strand
1
10.120183
Gyldenstens
155
156
ZMUC C-F- 123198
1
Fyn
4988640392
2013-04-27
ZMUC
JP
Denmark
JP
55.048084
Nyord
1
12.206333
155
156
ZMUC C-F-123325
1
Lolland, Falster, Mon
4988641649
2018-01-28
ZMUC
J. SOgaard Hansen
Denmark
J. Sogaard Hansen
55.985283
Auderod Skov
1
12.092983
155
156
ZMUC C-F-123504
1
Nordostsjaelland
4988640794
2016-05-26
ZMUC
JP
Denmark
JP
55.700718
Degnemosen
1
12.505867
Bronshoj
155
156
ZMUC C-F-122768
1
Nordostsjaelland
4988640523
[207,1399,1997,2025]
2013-11-16
ZMUC
JP
Denmark
JP
55.842335
Faergelunden
1
12.028916
155
156
ZMUC C-F-123084
1
Nordostsjaelland
4988641859
2013-05-15
ZMUC
JP
Denmark
JP
55.79563
Olstykke
1
12.170883
Lyngen
156
157
ZMUC C-F-123357
1
Nordostsjaelland
4988641181
2018-04-15
ZMUC
JP
Denmark
JP
55.562065
Kongelundsfortet
1
12.562183
Ud
156
157
ZMUC C-F-123568
1
Nordostsjaelland
4988640636
2015-01-19
ZMUC
JP
Denmark
Nord
JP
56.500816
Vinderup
1
8.7564
Landting
156
157
ZMUC C-F-123052
1
Nordvestjylland
4988641402
2013-05-31
ZMUC
JP
Denmark
JP
55.13875
Lakolk
1
8.493633
156
157
ZMUC C-F-122771
1
Sydjylland
4988640749
2019-02-17
ZMUC
JP
Denmark
JP
55.1449
Even Bro
1
12.010016
156
157
ZMUC C-F-123905
1
Sydsjaelland
4988640746
2019-04-25
ZMUC
JP
Denmark
JP
54.99575
Gammel Kalvehave
1
12.1372
156
157
ZMUC C-F- 123990
1
Sydsjaelland
4988641593
2012-01-14
ZMUC
JP
Denmark
JP
54.98725
Langebaek
1
12.104667
Osterskov
156
157
ZMUC C-F-123029
1
Sydsjaelland
4988641668
2019-01-03
ZMUC
JP
Denmark
JP
54.992867
Langebaek
1
12.111917
Osterskov
156
157
ZMUC C-F-123803
1
Sydsjaelland
4988641694
On Agonum versutum Sturm, 1824
2014-04-25
ZMUC
JP
Denmark
JP
55.072918
Ronne Plantage
1
14.7516
Skelsmyre
156
157
ZMUC C-F-122988
1
Bornholm
4988641690
On Agonum viduum (Panzer, 1797)
2017-05-07
ZMUC
JP
Denmark
JP
54.78285
Maribo
1
11.48265
Lysemose
156
157
ZMUC C-F-123148
1
Lolland, Falster, Mon
4988641590
On Anchomenus dorsalis (Pontoppidan, 1763)
2014-04-25
ZMUC
JP
Denmark
JP
55.160316
Osterlars
1
14.99605
Ost
156
157
ZMUC C-F-122689
1
Bornholm
4988641697
2016-04-22
ZMUC
JP
Denmark
JP
55.65148
Brondby
1
12.408116
156
157
ZMUC C-F-122520
1
Nordostsjaelland
4988640964
2013-09-21
ZMUC
JP
Denmark
JP
55.341167
Holtug Kalkbrud
1
12.4446335
Holtug Kalkbrud
156
157
ZMUC C-F-122829
1
Sydsjaelland
4988640473
On Anisodactylus binotatus (Fabricius, 1787)
2016-06-03
ZMUC
H. S. Pedersen
Denmark
JP
55.7119
Utterslev Mose
1
12.49845
156
157
ZMUC C-F-122589
1
Nordostsjaelland
4988640769
On Harpalus affinis (Schrank, 1781)
2014-10-16
ZMUC
JP
Denmark
JP
56.288033
Kjellerup
1
9.436617
156
157
ZMUC C-F-122798
1
Ostjylland
4988640596
2011-12-02
ZMUC
JP
Denmark
JP
55.81795
Naerum
1
12.544766
156
157
ZMUC C-F-122960
1
Nordostsjaelland
4988642118
On Loricera pilicornis (Fabricius, 1775)
2013-04-27
ZMUC
JP
Denmark
JP
55.048084
Lolland, Falster, Mon
1
12.206333
156
157
ZMUC C-F-123321
1
Lolland, Falster, Mon
4988642323
2016-05-26
ZMUC
JP
Denmark
JP
55.700718
Nordostsjaelland
1
12.505867
Bronshoj
156
157
ZMUC C-F-122767
1
Nordostsjaelland
4988640447
2017-04-14
ZMUC
JP
Denmark
JP
56.0245
Gurre Vang
1
12.491433
156
157
ZMUC C-F-122744
1
Nordostsjaelland
4988640471
2009-07-25
ZMUC
JP
Denmark
JP
55.432617
Soro
1
11.574616
Flommen
156
157
ZMUC C-F-122552
1
Sydsjaelland
4988641922
2013-07-22
ZMUC
H. Enghoff
Denmark
JP
55.418583
Kristiansminde
1
11.586817
Soro
156
157
ZMUC C-F- 122787
1
Sydsjaelland
4988641814
On Nebria brevicollis (Fabricius, 1792)
2006-05-01
2006-05-12
2006-05-01
ZMUC
J. HOier
Denmark
P. Jorum
56.048965
Nordostsjaelland
1
12.369017
156
157
ZMUC C-F-124332
1
Nordostsjaelland
4988640498
2006-05-01
2006-05-12
2006-05-01
ZMUC
J. HOier
Denmark
P. Jorum
55.7661
Harrevads Bro
1
12.376683
Lille Hareskov
156
157
ZMUC C-F-124334
1
Nordostsjaelland
4988640669
2006-09-01
2006-09-12
2006-09-01
ZMUC
J. HOier
Denmark
P. Jorum
55.79775
Norreskoven
1
12.384233
156
157
ZMUC C-F-124333
1
Nordostsjaelland
4988642060
On Oxypselaphus obscurus (Herbst, 1784)
2014-03-21
ZMUC
JP
Denmark
JP
56.018032
Hampen So
1
9.385817
157
158
ZMUC C-F-122918
1
Ostjylland
4988641479
2018-04-19
ZMUC
JP
Denmark
JP
55.557915
Rishave
1
10.400217
157
158
ZMUC C-F-123587
1
Fyn
4988640358
1993-07-08
ZMUC
S. Langemark & O. E. Meyer
Denmark
JP
54.773468
Krenkerup Haveskov
1
11.665916
157
158
ZMUC C-F-123405
1
Lolland, Falster, Mon
4988641193
1993-07-19
ZMUC
S. Langemark & O. E. Meyer
Denmark
JP
54.773468
Krenkerup Haveskov
1
11.665916
157
158
ZMUC C-F-12342
1
Lolland, Falster, Mon
4988640422
1993-08-16
1993-08-24
1993-08-16
ZMUC
S. Langemark & O. E. Meyer
Denmark
JP
54.773468
Krenkerup Haveskov
1
11.665916
157
158
ZMUC C-F-123410
1
Lolland, Falster, Mon
4988641039
2017-05-07
ZMUC
JP
Denmark
JP
54.78285
Maribo
1
11.48265
Lysemose
157
158
ZMUC C-F-123142
1
Lolland, Falster, Mon
4988642087
2018-06-04
ZMUC
JP
Denmark
JP
54.78285
Maribo
1
11.48265
Lysemose
157
158
ZMUC C-F-123648
1
Lolland, Falster, Mon
4988642150
2013-04-27
ZMUC
JP
Denmark
JP
55.048084
Nyord
1
12.206333
157
158
ZMUC C-F-123323
1
Lolland, Falster, Mon
4988642063
2018-05-26
ZMUC
JP
Denmark
JP
55.043934
Ulvshale
1
12.260633
157
158
ZMUC C-F-123627
1
Lolland, Falster, Mon
4988642077
2019-01-03
ZMUC
JP
Denmark
JP
54.838383
Bandholm
1
11.4675
Vest
157
158
ZMUC C-F-123844
1
Lolland, Falster, Mon
4988642071
2017-04-14
ZMUC
H. Enghoff
Denmark
JP
56.0245
Gurre Vang
1
12.491433
157
158
ZMUC C-F-122746
1
Nordostsjaelland
4988641712
2018-04-21
ZMUC
L. Thomas
Denmark
JP
55.821
Haettingsholm
1
12.232783
157
158
ZMUC C-F-123638
1
Nordostsjaelland
4988641878
2013-12-14
ZMUC
R. Aagaard Jensen
Denmark
JP
55.856735
Frederikssund
1
12.0589
Nord
157
158
ZMUC C-F-122891
1
Nordostsjaelland
4988641984
2013-04-14
ZMUC
JP
Denmark
JP
55.805534
Radvad
1
12.5523
Stampeskov
157
158
ZMUC C-F-123275
1
Nordostsjaelland
4988641434
2019-03-30
ZMUC
JP
Denmark
JP
55.83705
Trorod
1
12.51975
Vest
157
158
ZMUC C-F-123950
1
Nordostsjaelland
4988642226
2013-12-10
ZMUC
JP
Denmark
JP
55.74943
Tuse Naes
1
11.763617
Bognaes Skov
157
158
ZMUC C-F-123378
1
Nordvestsjaelland
4988641104
2017-02-18
ZMUC
JP
Denmark
JP
55.657818
Skarreso
1
11.385783
Nordbredden
157
158
ZMUC C-F-122818
1
Nordvestsjaelland
4988641751
2013-06-01
ZMUC
JP
Denmark
JP
55.016068
Draved Skov
1
8.97325
157
158
ZMUC C-F-122783
1
Sydjylland
4988642284
2012-01-14
ZMUC
JP
Denmark
JP
54.98725
Langebaek
1
12.104667
Osterskov
157
158
ZMUC C-F-123023
1
Sydjylland
4988642004
2013-06-02
ZMUC
JP
Denmark
JP
55.030483
Solsted Mose
1
8.839184
157
158
ZMUC C-F-122972
1
Sydjylland
4988640563
2018-03-24
ZMUC
JP
Denmark
JP
54.99575
Gammel Kalvehave
1
12.1372
157
158
ZMUC C-F-123511
1
Sydsjaelland
4988642155
[361,1058,1119,1146]
2018-04-27
ZMUC
JP
Denmark
JP
54.99575
Gammel Kalvehave
1
12.1372
157
158
ZMUC C-F-123600
1
Sydsjaelland
4988640387
2019-01-03
ZMUC
JP
Denmark
JP
54.992867
Langebaek
1
12.111917
Osterskov
157
158
ZMUC C-F-123808
1
Sydsjaelland
4988641337
2019-02-17
ZMUC
JP
Denmark
JP
55.220566
Sparresholm
1
11.957717
Storskov
157
158
ZMUC C-F-123929
1
Sydsjaelland
4988641101
On Paranchus albipes (Fabricius, 1796)
2018-06-24
ZMUC
JP
Denmark
JP
55.2276
Dondalens
1
14.884633
157
158
ZMUC C-F-123666
1
Bornholm
4988641403
2018-04-20
ZMUC
L. Kjaer-Thomsen
Denmark
JP
56.012585
Gribskov
1
12.367217
Kongens
157
158
ZMUC C-F-123636
1
Nordostsjaelland
4988642262
On Platynus assimilis (Paykull, 1790)
2017-05-26
ZMUC
L. K. Thomsen & L. Thomas
Denmark
JP
55.224815
Dondal
1
14.882267
157
158
ZMUC C-F-122911
1
Bornholm
4988640485
2018-06-23
ZMUC
JP
Denmark
JP
55.195667
Baggea
1
14.7543335
Svartingedalen
157
158
ZMUC C-F-123682
1
Bornholm
4988641871
2012-06-15
ZMUC
H. Liljehult
Denmark
JP
54.668568
Keldskov
1
11.55605
157
158
ZMUC C-F-122519
1
Lolland, Falster, Mon
4988640884
1993-05-05
ZMUC
S. Langemark & O. E. Meyer
Denmark
JP
54.773468
Krenkerup Haveskov
1
11.665916
157
158
ZMUC C-F-123393
1
Lolland, Falster, Mon
4988642125
[445,1399,1642,1669]
1993-05-12
ZMUC
S. Langemark & O. E. Meyer
Denmark
JP
54.773468
Krenkerup Haveskov
1
11.665916
157
158
ZMUC C-F-122579
1
Lolland, Falster, Mon
4988641467
[205,1154,1678,1705]
1993-05-25
ZMUC
S. Langemark & O. E. Meyer
Denmark
JP
54.773468
Krenkerup Haveskov
1
11.665916
157
158
ZMUC C-F-123407
1
Lolland, Falster, Mon
4988641858
1993-06-16
ZMUC
S. Langemark & O. E. Meyer
Denmark
JP
54.773468
Krenkerup Haveskov
1
11.665916
157
158
ZMUC C-F-122578
1
Lolland, Falster, Mon
4988640304
2018-05-26
ZMUC
JP
Denmark
JP
55.043934
Ulvshale
1
12.260633
157
158
ZMUC C-F-123626
1
Lolland, Falster, Mon
4988640795
2013-08-08
2013-08-22
2013-08-08
NEJ
I. Aggerholm
Denmark
JP
56.85875
Tofte Skov
1
10.251083
Engskoven
157
158
ZMUC C-F- 123435
1
Nordostjylland
4988641214
2018-05-20
ZMUC
L. Kjaer-Thsen
Denmark
L. Kjaer-Thomsen
55.614216
Lejre
1
11.943684
Herthadal
157
158
ZMUC C-F-123637
1
Nordostsjaelland
4988641114
2018-01-28
ZMUC
J. SOgaard Hansen
Denmark
J. Sogaard Hansen
55.680515
Lille Lyngby Mose
1
12.4848995
157
158
ZMUC C-F-123505
1
Nordostsjaelland
4988641763
2013-05-01
ZMUC
JP
Denmark
JP
55.593533
Tisso
1
11.307683
Nordostbredden
157
158
ZMUC C-F-123251
1
Nordvestsjaelland
4988640824
2013-05-01
ZMUC
JP
Denmark
JP
55.65708
Skarreso
1
11.3833
Nordbredden
157
158
ZMUC C-F-123350
1
Nordvestsjaelland
4988642335
2018-10-06
ZMUC
L. Kjaer-Thomsen & L. Thomas
Denmark
JP
158
159
55.43947
Ashoje Overdrev
1
12.105984
157
158
ZMUC C-F-123762
1
Sydsjaelland
4988641767
2018-03-27
ZMUC
JP
JP
54.98885
Langebaek
1
12.10595
Osterskov
158
159
ZMUC C-F- 123527
1
Ub
4988641928
[300,950,336,363]
2019-01-03
ZMUC
JP
JP
54.98885
Langebaek
1
12.10595
Osterskov
158
159
ZMUC C-F-123807
1
Ub
4988640659
2013-07-22
ZMUC
H. Enghoff
JP
55.418583
Kristiansminde
1
11.586817
Soro
158
159
ZMUC C-F-122786
1
Ub
4988640488
2014-07-02
ZMUC
JP
JP
55.418583
Kristiansminde
1
11.586817
Soro
158
159
ZMUC C-F-122865
1
Ub
4988641445
On Platynus livens (Gyllenhal, 1810)
2014-04-25
ZMUC
H. Liljehult
Denmark
JP
55.072918
Ronne Plantage
1
14.7516
Skelsmyre
158
159
ZMUC C-F-122712
1
Bornholm
4988641035
2019-03-31
ZMUC
JP
Denmark
JP
55.29475
Holmegards Mose
1
11.7988
158
159
ZMUC C-F-123964
1
Sydsjaelland
4988642198
On Pterostichus oblongopunctatus (Fabricius, 1787)
2006-07-01
2006-07-12
2006-07-01
ZMUC
J. HOier
Denmark
P. Jorum
55.757534
Store Sejben Mose
1
12.370283
Jonstrup Vang
158
159
ZMUC C-F-124336
1
Nordostsjaelland
4988642171
2006-05-01
2006-05-12
2006-05-01
ZMUC
J. HOier
Denmark
P. Jorum
55.7661
Harrevads Bro
1
12.376683
Lille Hareskov
158
159
ZMUC C-F-124335
1
Nordostsjaelland
4988640785
On Pterostichus vernalis (Panzer, 1796)
2014-04-25
ZMUC
JP
Denmark
JP
55.160316
Ost
1
14.99605
Osterlars
158
159
ZMUC C-F-122690
1
Bornholm
4988642181
2019-01-03
ZMUC
JP
Denmark
Mon
JP
54.838383
Vest
1
11.4675
Bandholm
158
159
ZMUC C-F-123865
1
Lolland, Falster, Mon
4988641495
2012-10-28
ZMUC
JP
Denmark
JP
55.7119
Utterslev Mose
1
12.49845
158
159
ZMUC C-F-122518
1
Nordostsjaelland
4988642152
2013-12-10
ZMUC
JP
Denmark
JP
55.74943
Bognaes Skov
1
11.763617
Tuse Naes
158
159
ZMUC C-F-123374
1
Nordvestsjaelland
4988641939
2014-01-26
ZMUC
JP
Denmark
JP
55.53275
Flasken
1
11.16385
Reerso
158
159
ZMUC C-F-123100
1
4988641528
2017-11-04
ZMUC
JP
Philippines
JP
55.969185
Dybeso
1
11.76035
158
159
ZMUC C-F-123210
1
4988641619
2017-02-18
ZMUC
JP
Philippines
JP
55.657818
Nordbredden
1
11.385783
Skarreso
158
159
ZMUC C-F-122817
1
4988640683
2018-03-24
ZMUC
JP
Philippines
JP
55.657818
Gammel Kalvehave
1
11.385783
Skarreso
158
159
ZMUC C-F-123508
1
Sydsjaelland
4988642248
[189,857,1248,1275]
2018-04-27
ZMUC
JP
Philippines
JP
54.99575
Gammel Kalvehave
1
12.1372
Skarreso
158
159
ZMUC C-F-123597
1
Sydsjaelland
4988641991
2019-04-25
ZMUC
JP
Philippines
JP
54.99575
Gammel Kalvehave
1
12.1372
Skarreso
158
159
ZMUC C-F-123992
1
Sydsjaelland
4988641257
2019-01-03
ZMUC
JP
Philippines
JP
54.992867
Osterskov
1
12.111917
Langebaek
158
159
ZMUC C-F-123822
1
4988640357
2019-04-25
ZMUC
JP
Philippines
JP
55.115215
Praesto
1
12.033916
158
159
ZMUC C-F-123982
1