Laboulbeniomycetes (Fungi, Ascomycota) of Denmark Santamaria, Sergi Pedersen, Jan European Journal of Taxonomy 2021 2021-11-26 781 1 425 B2MH7 Santam. & Cuesta-Segura [498,1089,264,291] Laboulbeniomycetes Laboulbeniaceae Rhachomyces Fungi Laboulbeniales 93 94 Ascomycota species spinosus  MB#834607  Fig. 18E–F    Nova Hedwigia110(3–4): 362 ( Santamaria et al.2020a).–    Type: “ Spain, León, Redipuertas, Valdelugueros, Riopinosvalley, 43°01′30″ N, 5°26′26″ W,  1346 ma.s.l.,  22.V.2016, A.D. Cuesta-Segura, on  Syntomus foveatus(Geoffroy)( Coleoptera Carabidae), det. A.D. Cuesta-Segura, slide BCB–SS2743n ( HOLOTYPE)”; BCB.    Diagnostic features Thallus small, up to 220 µm. Primary appendage three-celled. Primary septum constricted and darkened, with the dark brown pigmentation extending to the lower quarter of the basal cell of the primary appendage ( Fig. 18E, a). Second cell of the primary appendage bearing a stout spinous process on the margin ( Fig. 18E, sx). Apical cell of the primary appendage rounded distally, as long as the two lower cells together. Secondary axis three-celled, bearing five antheridial appendages.    Distribution and hosts On  Syntomus(Col. Carabidae). Only known from Spain(original description) and Belgium( Haelewaters & De Kesel 2020: pl. 71, as  Rhachomyces sciakyiW.Rossi).    Collections examined from Denmark  On  Syntomus foveatus(Geoffroy, 1785) (Col. Carabidae)   DENMARK–  Fyn(F)• Flyvesandetved Agernaes; 55°37.294′ N, 10°18.251′ E; NG86;  19 Apr. 2018;  JP1043; JPdet.; ZMUC C-F-123559. –   Lolland, Falster, Møn(LFM)• Ulvshale; 55°2.301′ N, 12°16.244′ E; UB20;  25 Sep. 2016;  JP144; JPdet.; ZMUC C-F-122623. –   Nordøstjylland(NEJ)• Sandmilen; 57°41.793′ N, 10°31.717′ E; NJ99;  21 Nov. 2017;  JP801; JPdet.; ZMUC C-F-123296. –   Nordøstsjaelland(NEZ)• Asserbo Plantage; 56°1.794′ N, 12°1.589′ E; UC11;  15 Oct. 2017;  JP576; JPdet.; ZMUC C-F-123062•  Liseleje; 56°1.048′ N, 11°58.454′ E; PH81;  30 Oct. 2016;  JP132; JPdet.; ZMUC C-F-122608. –   Nordvestsjaelland(NWZ)• Sandflugtsplantageved Rørvig; 55°57.126′ N, 11°43.060′ E; PH60;  4 Nov. 2017;  JP712; JPdet.; ZMUC C-F-123203.    Remarks First record from Denmark, third record after the original description. This species is related to  R. sciakyiW.Rossiand  R. lavagnei(F.Picard) W.Rossibut may be readily separated from them by the spinous process seen on the second cell of the primary appendage ( Santamaria et al.2020a). 4988641842 on Syntomus foveatus (Geoffroy) 2016-05-22 BCB A. D. Cuesta- Segura Spain A. D. Cuesta-Segura 1346 43.025 Redipuertas 19 -5.4405556 Leon 93 94 1 holotype 4988642093 2018-04-19 ZMUC JP Denmark JP 55.621567 Flyvesandet 1 10.304183 Agernaes 93 94 ZMUC C-F-123559 1 Fyn 4988642309 2016-09-25 ZMUC JP JP 55.621567 Ulvshale 1 10.304183 Agernaes 93 94 ZMUC C-F-122623 1 Lolland, Falster, Mon 4988642339 2017-11-21 ZMUC JP JP 55.621567 Sandmilen 1 10.304183 Agernaes 93 94 ZMUC C-F-123296 1 Nordostjylland 4988641667 2017-10-15 ZMUC JP JP 55.621567 Asserbo Plantage 1 10.304183 Agernaes 93 94 ZMUC C-F-123062 1 Nordostsjaelland 4988641359 2016-10-30 ZMUC JP Philippines JP 56.017467 Liseleje 1 11.974234 93 94 ZMUC C-F-122608 1 4988642347 2017-11-04 ZMUC JP Philippines JP 56.017467 Sandflugtsplantage 1 11.974234 Rorvig 93 94 ZMUC C-F-123203 1 Nordvestsjaelland