A revision of the Solomonius-group of the stinkbug tribe Halyini (Hemiptera: Pentatomidae: Pentatominae) WALL, MICHAEL A. Zootaxa 2007 2007-07-30 1539 1 1 84 (Horvath, 1915) Published 2007 Horvath 1915 [151,572,711,737] Insecta Brizica GBIF Animalia Hemiptera 37 38 Arthropoda species humeralis comb. nov.  ( Figs. 16D, 18J-L, 20C, 22A-B, 23) 1915  Neoptolemus humeralisHorváth, 244–245, figs. 11a, b. (keyed)  Diagnosis.Distinguished by relatively short labium, profemora armed with 7+ ventral barbs (distal two distinctly larger than proximal members of series), proctiger of male broadly inflated distally ( Fig. 18J).  Redescription. Body 2X longer than broad ( Fig. 16D). Vertex of head slightly convex, punctate. Juga apically pointed, swollen laterally. Juga and tylus subequal in length, depressed relative to remainder of head. First antennal segment surpassing apex of head. Eyes stalked; posterior margins converging with head at right angle. Pronotum rugose; anterolateral margins crenulate; lateral angles acute. Scutellum swollen anteriorly; apex broadly rounded, somewhat angulate, dorsally reflexed. Laterotergites with posterior angle produced into small thorn-like tubercles. Bucculae produced, becoming obsolete prior to reaching posterior margin of head; anterior angle acute. Labium reaching abdominal third sternite (second visible); ratios 1:1.75:1.5:1. Peritreme 1/4 as long as distance from ostiole of metathoracic scent gland to lateral margin of metapleuron. Episternal ridge located 2/3 of distance between ostiole of metathoracic scent gland and lateral margin of metapleuron. Venter of abdomen primarily convex; median flattened to slightly sulcate. Profemora with 7–8 ventral thorn–shaped spines, spines becoming larger on distal portions.  Color and vestiture.Ground color straminous to testaceous with black shining punctures. Antennal segments I–III testaceous with piceous markings; segments IV–V piceous. Legs mostly straminous, tibiae with two castaneous bands on longitudinal ridges. Abdomen mostly straminous; lateral portions of abdominal second sternite (first visible) with dark metallic area surrounding spiracles; remainder of spiracles black; intersegmental sutures laterally black. Antennal segments I–II and proximal portion of segment III shiny with sparse vestiture, remainder of segment III and segments IV–V with dense short pile.  Male.Ventral rim of pygophore slightly emarginate; dorsal rim produced as short lateral lobes ( Fig. 18J– L). Proctiger elongate, broadly dilated distally, distal portion wider than total length of proctiger; ventral surface sclerotized with microsculptured ridges ( Fig. 22); basal sclerite with posterior and anterior margins roughly straight ( Fig. 18L).  Female.Eighth gonocoxa triangular, narrowly acute and falcate apically. Ninth laterotergites swollen, almost cylindrical in cross section, narrowed apically ( Fig. 20C).  Variation.Specimens vary in ground coloration from straminous to testaceous to olivaceous.   FIGURE 22.Ventral view of male proctiger of  Brizica humeralis(Horváth)(A, scale = 200µm) and associated microridging (B, scale = 20µm).  Measurements.Males (n=4): body length 16.9 SD 0.7 (16.1–17.8), anterior pronotal width 3.0 SD 0.2 (2.8–3.3), humeral width 9.5 SD 0.5 (8.8–9.8), scutellar length 6.5 SD 0.2 (6.2–6.7), medial length of sixth sternite (fifth visible) 1.2 SD <0.1 (1.1–1.2), medial length of seventh sternite (sixth visible) 2.6 SD 0.05 (2.6– 2.7). Females (n=5): body length 17.9 SD 0.6 (17–18.7), anterior pronotal width 3.0 SD 0.3 (2.6–3.4), humeral width 9.3 SD 0.6 (8.3–9.8), scutellar length 6.5 SD 0.5 (5.9–6.8), medial length of sixth sternite (fifth visible) 3.9 SD 5.5 (1.4–13.9), medial length of seventh sternite (sixth visible) 2.6 SD 0.1 (2.4–2.8). Appendages in Table 3.   Specimensexamined.  PAPUA NEW GUINEA: Central Province, Daradae, nr. Javarere, Musgorve R., elev.  100 m, [ 9° 27' S, 147° 37’ E],  3 October 1958, coll. J.L. Gressitt– sex not recorded ( BPBM);  Koitaki, elev.  1000 m, [ 9° 22' S, 147° 26’ E],  1 October - 30 November 1928- sex not recorded ( BPBM);  Laloki, nr. Port Moresby, [ 9° 23' S, 147° 18’ E],  30 August - 2 September 1959, coll. T. Maa- sex not recorded ( BPBM);  Owen Stanley Range; Goilala: Loloipa, [ 8° 26' S, 146° 56’ E],  1–15 January 1958, coll. W. W. Brandt- sex not recorded ( BPBM);  Rigo District, [ 9° 48' S, 147° 33’ E],  1–30 September 1928- sex not recorded ( BPBM); SE: Brown River, [ 9° 15' S, 147° 4’ E],  12–23 October 1968, coll. Tawi Mena- sex not recorded ( BPBM); SE: Vanapa River, [ 9° 5' S, 146° 58’ E],  29 February 1960, coll. J. Sedlacek- sex not recorded ( BPBM);  St. Joseph’s River, [ 8° 50' S, 146° 34’ E], coll. W. MacGregor- 2 ♀♀( AM);   Eastern Highlands, Kundiawa, [ 6° 1' S, 144° 58’ E],  3 December 1964-  1 January 1965, coll. J. Sedlacek- ( BPBM); NE: Karimui, South of Goroka, elev.  1000–1080 m, [ 6° 30' S, 144° 51’ E],  7 June 1961, coll. J.L. Gressitt- sex not recorded ( BPBM); same locality,  13 August 1963, coll. J. Sedlacek- sex not recorded ( BPBM);   Milne Bay Province, Milne Bay, 10° 22' S, 150° 30’ E],  14–23 February 1969, coll. J. Sedlacek- sex not recorded ( BPBM);   Morobe Province, Aseki, elev.  1200–1300 m, [ 7° 21' S, 146° 53’ E],  1–28 February 1972- 1 ♂( UQIC);  Garaina, elev.  800 m, [ 7° 52' S, 147° 7’ E],  15 January 1968, coll. J. Sedlacek- sex not recorded ( BPBM);  Mt. Missim, elev.  1050–1100 m, [ 7° 13' S, 146° 48’ E],  1 October 1964, coll. J. Sedlacek- sex not recorded ( BPBM);  Wau Ecology Institute, elev.  1100–1300 m, [ 7° 20' S, 146° 43’ E],  1 January 1961-  31 December 1974- sex not recorded ( BPBM); same locality,  23 January 1974, coll. Bishop Acc1982.491 - 1 ♀( BPBM);   Northern Province, Mt. Lamington, [ 8° 56' S, 148° 10’ E],  1 January - 28 February 1929, coll. C, McNamara- 5 ♀♀, 3 ♂♂( AM); same locality,  1–30 June 1966, coll. S. Lippert- sex not recorded ( BPBM);  Sangara, [ 8° 48' S, 148° 14’ E],  31 August - 2 September 1963, coll. J. Sedlacek- sex not recorded ( BPBM).  Distribution.Primarily confined to the Papuan peninsula ( Fig. 23). 1958-10-03 BPBM Javarere & Musgorve R. & J. L. Gressitt Papua New Guinea 100 -9.45 Daradae 1299 147.61667 Specimens 38 39 1 Central Province 1928-10-01 1928-11-30 1928-10-01 BPBM Papua New Guinea 1000 -9.366667 Koitaki 1300 147.43333 38 39 1 Central Province 1959-08-30 1959-09-02 1959-08-30 BPBM T. Maa Papua New Guinea -9.383333 Laloki 1300 147.3 38 39 1 National Capital 1958-01-01 1958-01-15 1958-01-01 BPBM W. W. Brandt Papua New Guinea Owen Stanley Range -8.433333 Loloipa 1301 146.93333 Goilala 38 39 1 National Capital 1928-09-01 1928-09-30 1928-09-01 BPBM Papua New Guinea -9.8 Rigo District 1299 147.55 38 39 1 National Capital 1968-10-12 1968-10-23 1968-10-12 BPBM Tawi Mena Papua New Guinea -9.25 Brown River 1300 147.06667 38 39 1 National Capital 1960-02-29 BPBM J. Sedlacek Papua New Guinea -9.083333 Vanapa River 1300 146.96666 38 39 1 National Capital [269,1180,1352,1378] AM W. MacGregor Papua New Guinea -8.833333 St. Joseph's River 1300 146.56667 38 39 2 2 National Capital 1964-12-03 1965-01-01 1964-12-03 BPBM J. Sedlacek Papua New Guinea -6.016667 Kundiawa 1305 144.96666 38 39 1 Eastern Highlands 1961-06-07 BPBM J. L. Gressitt Papua New Guinea 1040 -6.5 South of Goroka 1304 144.85 Karimui 38 39 1 Eastern Highlands [380,1291,1472,1498] 1963-08-13 BPBM J. Sedlacek Papua New Guinea 1040 -6.5 South of Goroka 1304 144.85 Karimui 38 39 1 Eastern Highlands 1969-02-14 1969-02-23 1969-02-14 BPBM J. Sedlacek Papua New Guinea -10.366667 Milne Bay Province 1298 150.5 38 39 1 Milne Bay 1972-02-01 1972-02-28 1972-02-01 UQIC Papua New Guinea 1250 -7.35 Aseki 1303 146.88333 38 39 1 1 Morobe 1968-01-15 BPBM J. Sedlacek Papua New Guinea 800 -7.866667 Garaina 1302 147.11667 38 39 1 Morobe 1964-10-01 BPBM J. Sedlacek Papua New Guinea 1075 -7.2166667 Mt. Missim 1303 146.8 38 39 1 Morobe 1961-01-01 1974-12-31 1961-01-01 BPBM Papua New Guinea 1200 -7.3333335 Wau Ecology Institute 1303 146.71666 38 39 1 Morobe 1974-01-23 1974-12-31 1961-01-01 BPBM Bishop Acc Papua New Guinea 1200 -7.3333335 Wau Ecology Institute 1303 146.71666 38 39 1 1 Morobe 1929-01-01 1929-02-28 1929-01-01 AM McNamara Papua New Guinea -8.933333 Mt. Lamington 1300 148.16667 38 39 8 5 3 Northern [538,1431,1792,1818] 1966-06-01 1966-06-30 1966-06-01 BPBM S. Lippert Papua New Guinea -8.933333 Mt. Lamington 1300 148.16667 38 39 1 Northern [151,1428,1832,1858] 1963-08-31 1963-09-02 1963-08-31 BPBM J. Sedlacek Papua New Guinea -8.8 Sangara 1300 148.23334 38 39 1 Northern