Phanaeus huichol Moctezuma et al ., 2017: 123 . Phanaeus huichol Kohlmann et al . 2018: 69 Moctezuma et al . 2019: 252–253 Phanaeus ( Notiophanaeus ) huichol Kohlmann et al . 2018: 81 Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species Moctezuma, Victor Halffter, Gonzalo European Journal of Taxonomy 2021 2021-04-28 747 1 71   Phanaeus huichol  Moctezuma et al ., 2017: 123  .  Kohlmann et al . 2018: 69   Kohlmann et al . 2018: 81  4FSJP Moctezuma, Sanchez-Huerta & Halffter, 2017 Moctezuma, Sanchez-Huerta & Halffter 2017 [408,1179,1783,1810] Insecta Scarabaeidae Phanaeus Animalia Coleoptera 21 22 Arthropoda species huichol   Figs 1G, 15, 18H, 19H      Phanaeus huichol Moctezuma et al., 2017: 123.    Phanaeus huichol–  Kohlmann et al. 2018: 69, 81, 83, 88. —  Moctezuma et al. 2019: 252–253, fig. 5.  Phanaeus( Notiophanaeus) huichol–  Kohlmann et al. 2018: 81.    Type material   Holotype   MEXICO• ♂,  Moctezuma et al. 2017: 123(revised); Jalisco, Mazamitla; TAMU.  Non-type material revised( 18 ♂♂, 20 ♀♀)   MEXICO–  Jalisco• 1 ♀; “ Sierra de Talpa(Los Venados CT).  12–15-VIII- 2017. 1570 m. G. Nogueira Col.”; GHC•  2 ♂♂, 1 ♀; “ Sierra de Talpa. 20°22′35.29″ N, 104°49′8.87″ O,  14/VIII/ 2015. 1380 m. G. Nogueira Col.”; VMC•  2 ♂♂, 1 ♀; “ Sierra de Talpa. 20°22′35.29″ N, 104°49′8.87″ O,  22–26/ VIII/ 2013. 1595 m. G. Nogueira Col.”; VMC•  2 ♂♂, 1 ♀; “ Tequila, Volcán de Tequila. 20°47′18.03″ N, 103°50′28.99″ O,  25/VIII/ 2012. 1890 m. G. Nogueira Col.”; VMC•  2 ♂♂, 1 ♀; “ Tequila, Volcán de Tequila. 20°47′18.03″ N, 103°50′28.99″ O,  24–27/VIII/ 2012. 1665 m. (CT). G. Nogueira Col.”; VMC•  1 ♂, 2 ♀♀; “ Telcome, Yolosta. 20°10′40.113″ N, 103°41′25.35″ O,  2/VII/2017. D  1–2. 1845m. G. Nogueira Col.”; VMC•  4 ♂♂, 4 ♀♀; “ El Jacal,  1830 m,  12–14/VIII/94. G. Nogueiracol.”; IEXA•  1 ♂, 2 ♀♀; “S. Manatlán, 13/ 16–IX– 95, 1650 m, G. Nogueiracol.”; IEXA•  3 ♂♂, 2 ♀♀; “ S. Manatlán, 18/ 20–IX–1995,  1400 m, G. Nogueiracol.”; IEXA•  1 ♂, 4 ♀♀; “ S. Manatlán, 18/ 20–IX– 1995, 1650 m, G. Nogueiracol.”; IEXA. –   Michoacán• 1 ♀; “  12 kmS Uruapan, Cascada Tzararacua.  1450 m.  21-VIII-97horse dung”; VMC.   Type locality  Mexico, Jalisco, Mazamitla.    Distribution Sierra Madre Occidental; Jalisco, northern Michoacán, Nayaritand southern Sinaloa( Fig. 15). The first record of  P. huicholfrom Michoacánis presented herein.    Remarks The original description of  P. huicholindicates that the colour pattern varies from dark metallic green, bright metallic green, to dark green with blue sheen. After revising additional specimens, a bright metallic green morph with red sheen was found by us. To the original description we add the elytral striae impressed basally as distinct fossae; and central ridge and column of endophallite copulatrix similar in size ( Fig. 1G). For the female, the head showing a cephalic trituberculate carina with middle tubercle more frontally projected, carinate and less prominent than lateral tubercles; frons with almost effaced punctures; pronotal sculpture completely smooth, with almost effaced punctures; pronotal process trituberculate, with posterior concavity, followed posteriorly by reduced, rounded tubercle; middle tubercle dentiform or rounded, more developed and projected frontally than lateral tubercles; lateral tubercles rounded or carinate; posterior pronotal midline completely effaced. 3093980606 [189,1150,349,376] TAMU Mexico Mazamitla Moctezuma 22 23 1 1 Jalisco holotype 3093980381 2017-08-12 2017-08-15 2017-08-12 GHC G. Nogueira Col. Mexico Sierra de Talpa 22 23 1 1 Jalisco 3093980494 2015-08-14 VMC G. Nogueira Col. Mexico 20.376469 Sierra de Talpa. 1 104.81913 22 23 3 1 2 Jalisco 3093980568 2013-08-22 2013-08-26 2013-08-22 VMC G. Nogueira Col. Mexico 20.376469 Sierra de Talpa. 1 104.81913 22 23 3 1 2 Jalisco 3093980512 2012-08-25 VMC G. Nogueira Col. Mexico 20.788342 Volcan de Tequila. 1 103.841385 Tequila 22 23 3 1 2 Jalisco 3093980621 2012-08-24 2012-08-27 2012-08-24 VMC G. Nogueira Col. Mexico 20.788342 Volcan de Tequila. 1 103.841385 Tequila 22 23 3 1 2 Jalisco 3093980303 2017-07-02 VMC G. Nogueira Col. Mexico 1845 20.177809 Yolosta. 1 103.69038 Telcome 22 23 3 2 1 Jalisco 3093980435 [513,1382,719,746] 1994-08-12 1994-08-14 1994-08-12 IEXA G. Nogueira Mexico 1830 El Jacal 22 23 8 4 4 Jalisco 3093980616 [189,1057,756,783] 1995-09-13 IEXA G. Nogueira Mexico 1650 Manatlan 22 23 3 2 1 Jalisco 3093980491 1995-09-20 IEXA G. Nogueira Mexico 1400 S. Manatlan 22 23 5 2 3 Jalisco 3093980484 1995-09-20 IEXA G. Nogueira Mexico 1650 S. Manatlan 22 23 5 4 1 Jalisco 3093980361 1997-08-21 VMC Mexico 1450 Cascada Tzararacua. 22 23 1 1 Michoacan