Omalium caesum Gravenhorst, 1806: 209 Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae, Omaliini) Shavrin, Alexey V. Khachikov, Eduard A. Zootaxa 2023 2023-01-09 5227 4 443 458 499S8 Gravenhorst, 1806 Gravenhorst 1806 [151,605,1806,1833] Insecta Staphylinidae Omalium Animalia Coleoptera 1 444 Arthropoda species caesum  ( Figs. 1, 11–12, 33)      Omalium caesum Gravenhorst, 1806: 209   Material examined.  RUSSIA: ROSTOVAReA:  1 ♀: Sholokhovskiy District, Elanskaya St.24–  26.06.1999. E.A. Khachikovleg. (cKh);  1 ♀: same data, Veshenskaya.  05.09.2002(cKh);  1 ♀: same data.  26.06.2004(cKh); VOLGOGRAD AReA:1 ♁: Elton Lake. 01– 10.08.2006. K.V. Makarov & A.V. Matalin leg. ( MSPU); KRASNODAR TeRRITORY: 1 ♀: Kustshevskiy District, Pervomaiskiy. Forest belt, sieve+sifter. 04.06.2009. L.B. Rybalov leg. ( ZMM); 1 ♀: Ubinskaya. 25.05.1984. I.A. Ushakov leg. ( ZMM); 1 ♀: Temnolesskaya. 25.06.1999. V.V. Grebennikov leg. (cSh); STAVROPOL TeRRITORY: 1 ♀: Pyatigorsk, Mashuk Mt. 17.06.1993. E.A. Khachikov leg. (cKh); 1 ♀: Essentuki. 10.03.2000. E.A. Khachikov leg. (cKh); 1 ♀: Zheleznovodsk, Razvalka. 08.06.1985. A.B. Ryvkin leg. (cR); ADYGeA:2 ♁♁: Maykop. 06.10.2009. A.R. Bibin leg. (cKh); 1 ♁, 1 ♀: Maykop, Khanskaya, right bank of Belaya River, 44°41′03.54′′N 39°55′41.25′′E. Flood-plain forest. 15.05.2014. E.A. Khachikov leg. (cKh); 1 ♁: same data, 04.06– 02.07.2009(cKh); 4 ♁♁, 6 ♀♀: Khanskaya. 19.06.2015. E.A. Khachikov leg. (cSh); 2 ♁♁, 3 ♀♀: same data. 16.05.2015(cKh, cSh); 1 ♀: Mezmay. 19– 25.06.1996. E.A. Khachikov leg. (cKh) 1 ♁: Caucasus Nature Reserve, Guzeripl. 21.06.1986. I.A. Ushakov leg. ( ZMM); 2 ♁♁, 2 ♀♀: same data, 18– 23.05.2015. E.A. Khachikov leg. (cSh); 3 ♁♁, 1 ♀: same data, 22– 23.08.2015(cKh, cSh); 5 ♁♁, 2 ♀♀: 05.10.2015(cKh, cSh); 2 ♁♁, 5 ♀♀: same data, Abago (cKh, cSh); 1 ♁, 1 ♀: Abago. 1000 ma.s.l. Forest with  Fagus,  Abies, motley grass. 02– 20.05.2009. S. Chumachenko leg. (cKh); 3 ♁♁: Abago, Klenovaya Roshcha. 1750 ma.s.l. 29.08– 28.09.2009. S. Chumachenko leg. (cKh); 1 ♀: 11 kmGuzeripl-Abago road. 27.07.2014. E.A. Khachikov leg. (cKh); 4 ♁♁, 2 ♀♀: same data. 2015 (cSh); KARACHAY-CHeRKeSSIA:1 ♁, 1 ♀: Teberda, Dombay. 07.2010. E.A. Khachikov leg. (cSh); 1 ♀: Teberda State Nature Biosphere Reserve,Alibek Gorge, Dvuyazychniy Glacier. 12.07.1996. E.A. Khachikov leg. (cKh); 1 ♀: same Nature Reserve, Malaya Khatipara Mt. 1900 ma.s.l. 29.05.1985. A.B. Ryvkin leg. (cR); KABARDINO-BALKARIA:1 ♁, 1 ♀: Kabardino-Balkaria, near Elbrus Mt. 2100–2350 ma.s.l.,  Betulaand  Pinusforest. 15– 19.08.1989. S.I. Golovatch leg. (cSh); NORTH OSSeTIA-ALANIA: 1 ♀: bank of Terek River near Elkhotovo. 29.07.1984. S.K. Alexeev leg. ( ZMM); 63 ♁♁, 43 ♀♀: Ardon River, Zincar vill. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); 6 ♁♁, 6 ♀♀: Ardon River, Zincar.  Taxus, traps. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); 16 ♁♁, 4 ♀♀: Biragzang, southern slope. Traps. 13.04.1984. S.K. Alexeev leg. (cSh, cR, ZMM); GEORGIA: SAMeGReLO-ZeMO SVANeTI: 1 ♀: 30 kmNE Khaishi, near Kherkhvashi. 1700 ma.s.l., litter near stream. 29.08.1986. A.B. Ryvkin leg. (cR); IMeReTI: 1 ♀: Mukhura, ca. 15 kmE of Tkibuli, 700–800 ma.s.l., forest with  Castanea,  Fagus,  Carpinusetc., under bark and stones. 07– 09.05.1987. S.I. Golovatch & K.Yu. Eskov leg. ( ZMM); MTSKHeTA-MTIANeTI:2 ♁♁: Saguramo Reserve, Zedazeni. 1100–1200 ma.s.l., forest with  Fagus,  Carpinus,  Aceretc,, litter and under bark. 20.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); TBILISI:9 ♁♁, 12 ♀♀: Tskhneti, 1150–1300 ma.s.l.,  Fagus,  Carpinus,  Aceretc. forest, litter. 16, 18.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cR, cSh); KVeMO-KARTLI: 2 ♀♀: Algeti National Park, W of Manglisi. 1400–1500 ma.s.l., forest with  Fagus,  Picea,  Abiesetc., in litter and under bark. 16– 18.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh); KAKHeTI:7 ♁♁, 3 ♀♀: Mariamjvari Nature Reserve, NNE of Sagarejo. 1150–1250 ma.s.l., forest with  Fagus,  Carpinus,  Acer,  Pinusetc. 13– 14.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); 1 ♁: N Kvareli. 700–750 ma.s.l., forest with  Fagus,  Carpinus,  Quercusetc., in litter and under bark. 04.05.1987. S.I. Golovatch & K.Yu. Eskov leg. ( ZMM); 1 ♀: Lagodekhi Nature Reserve. 500 ma.s.l. wet litter in forest near river. 05– 06.05.1983. S.I. Golovatch & A.B. Ryvkin leg. ( ZMM);  ARMENIA: LORI:1 ♁, 1 ♀: Odzun W of Alaverdi. 1500–1550 ma.s.l., forest with  Quercus,  Fagus,  Carpinus, etc., litter and under stones. 23– 24.07.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh);  TAVUSH: 1 ♀: Berdavan, ca 10 kmN of Noyemberyan. 900–950 ma.s.l. Forest with  Quercus,  Carpinus,  Aceretc., under stones. 24– 25.05.1987. S.I. Golovatch & K.Yu. Eskov leg. ( ZMM); 2 ♁♁, 1 ♀: Dilijan National Park, Akhnabat valley near Salakh. 1400–1500 ma.s.l., forest with  Fagus,  Taxus,  Carpinusetc., in litter, under bark and stones. 17.04.1983. S.I. Golovatch leg. (cSh, ZMM); 3 ♁♁, 3 ♀♀: W Shamshadyn halfway between Ijewan and Bered. 1500–1600 ma.s.l., forest with  Fagus,  Carpinus,  Aceretc. 26– 27.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM);  AZERBAIJAN: SHAKI-ZAQATALA:4 ♁♁, 2 ♀♀: N Vartashen (Oðuz). 900 ma.s.l, forest with  Fagusand  Quercus, in litter. 02.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh, ZMM); 1 ♁: NW above Baş Layisqi, ca. 20 kmNNW of Sheki. 1250 ma.s.l., forest with  Fagus,  Carpinus,  Aceretc., litter. 03.05.1987. S.I. Golovatch & K.Yu. Eskov leg. (cSh);  TURKEY: BURSA:1 ♁: Uludað. 1900 ma.s.l. 22.07.1969. C. Besuchet leg. ( MHNG); 1 ♁: Uludað, 17 kmBursa. 1100 ma.s.l. 13.05.1976. C. Besuchet & I. Löbl leg. ( MHNG);  ZONGULDAK:1 ♁, 1 ♀: 17 kmN Safranbolu. 1000 ma.s.l. 16.05.1976. C. Besuchet & I. Löbl leg. ( MHNG);  BOLU: 1 ♀: Abant. 1450 ma.s.l. 25.05.1967. C. Besuchet leg. ( MHNG);  MANISA: 2 ♀♀: Salihli. 500 ma.s.l. 29.04.1975. C. Besuchet & I. Löbl leg. ( MHNG);  GȔMȔşHANe: 2 ♀♀: Erzincan-Keklit. 2100 ma.sl. 04.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG);  ERZINCAN: 1 ♀: Tercan, Euphrate. 1400 ma.s.l. 06.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG);  ARTVIN:2 ♁♁: SW Artvin. 1900 ma.s.l. 09.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG); 2 ♀♀: S Pirnalli, Karkal Daði, 1250 ma.s.l. 11.06.1986. C. Besuchet, I. Löbl & D. Burckhardt leg. ( MHNG);  HATAY:57 ♁♁, 28 ♀♀: Antakya, Kişlak-Şenköy. 800–850 ma.s.l. 02.05.1978. C. Besuchet & I. Löbl leg. ( MHNG).   FIGURES 1–10.Habitus of  Omalium: 1—  O. caesum(Turkey, Bursa: Uludağ), 2—  O. cinnamomeum(Turkey, Antakya: KişlakŞenköy), 3—  O. excavatum(Russia, Karachay-Cherkessia: Arkhyz), 4—  O. falsum(Georgia, Tskhneti), 5—  O. littorale(Russia, Adygea: Abago), 6—  O. riparium impar(Russia, Krasnodar Territory: Khosta), 7—  O. rivulare(Armenia, Alaverdi), 8—  O. rugatum(Iran, Zagros Mts.), 9—  O. turcicum(Russia, Adygea: Lagonaki), 10—  O. wunderlei(Turkey, Gümüşhane: ErzincanKelkit). Scale bars: 1.0 mm.  Published records. AZERBAIJAN: Leder 1886: 120;  GEORGIA: Hochhuth 1849: 204, Zanetti 2002: 56; NAGORNO-KARABAKH: Assing & Schülke 2019: 99;  RUSSIA: ADYGeA: Zanetti 2002: 57;  DAGeSTAN: Khachikov et al. 2021: 250;  KABARDINO-BALKARIA: Khachikov 1998: 6;  KRASNODARTeRRITORY: Roubal 1911: 1; Zanetti 2002: 57;  TURKEY:J. Sahlberg 1913: 58; Bordoni 2010: 47.   Distribution.  Omalium caesumis widely distributed in the Palaearctic Region: Europe (including British islands), north Africa, Middle Asia, Western and Eastern Siberia ( Schülke & Smetana 2015).   Remarks.Habitus as in Fig. 1. Aedeagus as in Figs. 11–12. Female genital segments as in Fig. 33.  It is here recorded from several regions of SW Russia( StavropolTerritory, Karachay-Cherkessia, North Ossetia-Alania) and Armeniafor the first time. 4012666329 [395,682,1970,1996] Russia ROSTOV 1 444 1 Rostov 4012666308 1999-06-26 E. A. Khachikov Russia Elanskaya St. Sholokhovskiy District 1 444 1 1 Rostov 4012666323 [413,962,2006,2033] 2002-05-09 E. A. Khachikov Russia Veshenskaya. Sholokhovskiy District 1 444 1 1 Rostov 5891682301 2004-06-26 E. A. Khachikov Russia 2 445 Veshenskaya. Sholokhovskiy District 1 444 1 1 Rostov 4012666303 Russia Stavropol 4 447 1 Stavropol'