The Neotropical species of the genera Pseudespeson L, 1994 and Espeson S, 1882 (Coleoptera: Staphylinidae: Osoriinae) Irmler, U. Beiträge Zur Entomologie = Contributions to Entomology 2012 2012-12-20 62 2 331 360 3BJG6 Irmler, 2012 Irmler 2012 [265,432,175,199] Insecta Staphylinidae Espeson GBIF Animalia Coleoptera 6 337 Arthropoda species dybasi sp. nov.   Type material:   Holotype, male:  Panama: Chiriqui, Finca Palo Santo,  5.3.1959, leg. H. S. Dybas( AMNH).   Paratypes: 179 specimenswith same date as holotype; NW Volcán de Chiriqui, litter near stream ( 82°40'32"W, 8°49'19"N),  1450 melevation, 4 males, 6 females, 16.5.2001, 1 female, 17.5.2001, leg. L. Herman;  25.7 kmNW Volcán de Chiriqui, litter near stream ( 82°46'24"W, 8°50'06"N),  1150 melevation, 3 females, 27.12.2001, leg. L. Herman;   11.5 kmE Volcán de Chiriqui, Rio Sereno, tree fall litter in forest ( 82°46'24"W, 8°50'05"N),  1150 melevation, 1 female, 26.12.2001, leg. L. Herman;  Panama, road to Cerro Campana, litter near seep ( 79°54'50"W, 8°41'20"N),  645 melevation, 4 specimens, 1.1.2002, leg. L. Herman; road to Cerro Campana, litter near stream ( 82°55'12"W, 8°41',00"N),  730 melevation, 3 specimens, 22.12.2001, leg. L. Herman;   Costa Rica: Puntarenas,  35 kmNE San Vito, nr. Las Alturas( 82°50.08'W, 8°56.16'N), trail to Cerro Echandi,  1950 melevation, beyond trail marker 30, litter near stream, 78 specimens, 19.3.1991, leg. L. Herman;  7.5 kmon trail to Cerro Echandi, Rio Bella Vista, litter near stream, 34 specimens, 21.3.1991, leg. L. Herman;  Las Cruces Botanical Garden, near San Vito( 82°57.19W, 8°47.28'N),  1066 melevation, 1 specimen, 27.2.1985, leg. L. Herman;  Alajuela, nr. Poasito( 84°11.35'W, 10°10.09N),  6.6 kmW Jct. Rt.126(9) & 120, 2011 m elevation, VolcánPoas, litter near stream, 6 females, 16.3.1991, leg. L. Herman;  Cartago,  16.5 kmN Capellades( 83°47.07'W, 9°55.27'N), Rd.to Volcán de Turrialba,  2538 melevation, leaf litter under hummock in pasture, 7 specimens, 14.3.1991, leg. Herman;  Heredia,  2 kmS Sacramento, Rd.to VolcánBarba,  2011 melevation, litter near stream, 1 male, 15.3.1991, leg. L. Herman;  La Selva Biol. Station, 1 specimen, 18.2.1985, leg. L. Herman;   27 kmN Heredia( 83°59'W, 10°39'W), Rt9, 1828 m elevation, 1 specimen, 9.2.1985, leg. L. Herman;   Ecuador: Pichincha,  18-20 kmNE Alluriquin, old Quito-Sto Domingo rd.,  1460 melevation, litter, 1 specimen, 21.10.1988, leg. L. Herman; 1950 melevation, litter, 1 specimen, 23.10.1988;  43 kmN, Las Palmeras, old Qt Sto. Dgo. rd.km 59, litter, 2 females, 23.10.1988, leg. L. Herman;  Napo,  7 kmW Baeza, litter,  2400 melevation,  1 male,  31.10.1988, leg. L. Herman( AMNH, UIC).   Diagnosis: The species resembles  E. mexicanusand  E. simplexin size and colour. It can be easily differentiated from these two species by the elongate antennae. Whereas in  E. mexicanusand  E. simplexantennae are only slightly longer than head and pronotum combined, antennae of  E. dybasiare as long as head plus pronotum plus half of elytra. Moreover, the shape of the elytra also differs distinctly between  E. dybasiand both  E. mexicanusand  E. simplex, respectively. Width of elytra of  E. dybasiis more than 1.5 times as wide as pronotum, whereas it is only 1.3 times as wide in  E. mexicanusand  E. simplex. In all three species tergite VIII is slightly emarginate at posterior edge. In  E. mexicanusthe emargination is weak and smooth without distinct acute prominences at outer posterior angles. In  E. dybasia short stylus is placed at posterior angles, while an acute teeth is present in  E. simplex.   Description:  Length: 2.3 mm. Colour: brown; elytra and antennae red; legs yellow.  Head: 0.30 mmlong, 0.40 mmwide; eyes distinctly prominent; temples half as long as eyes; smoothly rounded to neck; fore-head distinctly narrowed to acute front edge of clypeus; densely and deeply punctate; distance between setiferous punctures on average less than 1/2 of diameter of punctures; vertex with impunctate midline; without microsculpture; surface shiny.  Antennaelonger than head and pronotum combined; 2 ndantennomere short; 3 rdconical antennomere longer than 2 nd; 3 rdto 7 thantennomeres longer than wide; 8 thantennomere slightly narrower and shorter than preceding and following antennomeres.  Pronotum: 0.38 mmlong, 0.38 mmwide; smoothly rounded in anterior half; distinctly emarginate in posterior half; lateral margin not visible in dorsal aspect; setiferous punctures dense and coarse; with narrow impunctate midline; on average distance between punctures half as wide as diameter of punctures; with small, but deep circular impression in central position of posterior half.  Elytra: 0.55 mmlong, 0.60 mmwide; sides smoothly rounded; widest in posterior third; setiferous punctation distinctly sparser and weaker than on pronotum; distance between punctures at least as wide as diameter of punctures, but on average distinctly wider.  Abdomenwith abdominal tergite III densely and deeply punctate; distance between punctures 1/2 as wide as diameter of punctures; density of punctation decreasing from tergite III to tergite VII; distance between punctures on tergite VII 2 to 3 times as wide as diameter of punctures; abdominal tergite VIII widely emarginate and with acute angles; on top of each acute angle a short stylus; at base of tergite VIII two setiferous pores within irregular dark spots.  Aedeaguswith more or less straight central lobe; paramera straight and with small irregular plate at apex.   Etymology: The specific name refers to the name of its first collector Dr. H. S. Dybas. 3118920365 [189,1136,274,298] 1959-03-05 AMNH H. S. Dybas Panama Finca Palo Santo 6 337 1 Chiriqui holotype 3118920341 1450 8.821944 Volcan de Chiriqui 21 -82.67555 6 337 179 paratype 3118920363 1150 8.835 Volcan de Chiriqui 21 -82.77334 6 337 1 paratype 3118920385 1150 8.834722 Rio Sereno 21 -82.77334 6 337 1 paratype 3118920339 645 8.688889 Cerro Campana 21 -79.91389 Panama 6 337 1 paratype 3118920343 730 Cerro Campana -82.92 6 337 1 paratype 3118920357 Cerro Echandi Costa Rica 1950 8.936 Las Alturas 13 -82.83466 6 337 1 Puntarenas paratype 3118920323 Costa Rica Rio Bella Vista Cerro Echandi 6 337 1 Puntarenas paratype 3118920301 Costa Rica 1066 8.788 San Vito 13 -82.95316 Las Cruces Botanical Garden 6 337 1 Puntarenas paratype 3118920315 Costa Rica 10.168167 Volcan 13 -84.18916 Poasito 6 337 1 Alajuela paratype 3118920336 Rd. Costa Rica 2538 9.921166 Volcan de Turrialba 13 -83.7845 6 337 1 Cartago paratype 3118920389 Costa Rica 2011 Volcan Rd. 6 337 1 Heredia paratype 3118920387 Costa Rica La Selva Biol. Station 6 337 1 Heredia paratype 3118920392 Rt Costa Rica 27 km N Heredia -10.65 6 337 1 Heredia paratype 3118920340 Ecuador 1460 Quito-Sto Domingo rd. 6 337 1 Pichincha paratype 3118920314 Ecuador Qt Sto. Dgo. rd. Las Palmeras 6 337 1 Pichincha paratype 3118920364 Ecuador 2400 7 km W Baeza 6 337 1 Napo paratype 3118920324 [443,986,1018,1042] 1988-10-31 AMNH, UIC L. Herman Ecuador 6 337 1 1 Napo paratype