Dyscritobaeus Dyscritobaeus sulawensis : Mineo, O’Connor et Ashe (2009) Dyscritobaeus sulawensis : O'Connor & Ashe (2012) Dyscritobaeus hannibal : O’Connor et Ashe (2011) Revision of Afrotropical Dyscritobaeus Perkins, 1910 (Hymenoptera: Scelionidae) Tortorici, Francesco Caleca, Virgilio Noort, Simon Van Masner, Lubomir Zootaxa 2016 4178 1 1 59 Mineo, O'Connor et Ashe Mineo, O'Connor et Ashe [151,788,583,610] Insecta Scelionidae Dyscritobaeus Animalia Hymenoptera 28 29 Arthropoda species sulawensis  urn:lsid:zoobank.org:act:2E29CC1E-9C7D-4E55-89A9- E75772A841E6 http://hol.osu.edu/index.html?id=259484   Mirotelenomussp.: Mineo & Villa (1982b), p. 138, Figs XXXV, XXXVI, XXXVII  Mirotelenomussp.: Mineo & Villa (1982a), p.178, Figs XXV, XLIII, XLVI, LV  Dyscritobaeussp. (a): Mineo & Caleca (1992) Fig. 1, p. 12; Fig. 3, p. 13.    Dyscritobaeussp. (a): Caleca & Mineo (1995)Figs III, 2, 3 p.12; Figs IV, 6, 7 p.13; Figs V, 3, 4 p.15; Fig. VII, p. 18.  Dyscritobaeus sulawensis: Mineo, O’Connor et Ashe (2009), p. 175, original description.  Dyscritobaeus sulawensis: O'Connor & Ashe (2012), p. 314.   Dyscritobaeus hannibal: O’Connor et Ashe (2011), p. 347. syn. nov.   Diagnosis.preoccipital area. The marginalis and stigmalis are normally developed as in  D. comitansand most of the other species in the genus; this character distinguishes this species from  D. armatus(Dodd), having a reduced stigmalis. T2 with a rounded-shaped specillum in females ( Fig. 11), but absent in males (Fig. 34); fore wing shape and angle between stigmalis and postmarginalis similar to  D. comitans; the combination of the presence of a specillum in females, absence of specillum in males and features of the fore wing is shared with  D. madagascarensis,but distinguishes this species from all species known to be close to  D. orientalis(1 described,  D. antanananrivensisTortorici et Caleca  sp. nov.and 9 undescribed species). In females the antennal segment A12 is shorter than A11 ( Fig. 17), with a corresponding claval sensilla formula: 0:2:2:2:1:0 (Fig. 18); this character state is unique in the genus. Adorsal tooth that is longer than the ventral tooth ( Fig. 74) distinguishes the species from  D. madagascarensisin both sexes. In females the specillum is rounded in comparison with drop-shaped specillum of  D. madagascarenis; in males the distance between the lateral ocellus and occipital carina is less than the ocellar diameter; the occipital carina is depressed at the vertex in comparison with  D. madagascarensis(in which the distance between the lateral ocellus and occipital carina is equal to the ocellar diameter; the occipital carina is less depressed at the vertex). After comparing the description of  D. sulawensis( Mineo et al.2009)and photographs of the holotypekindly taken by James O’Connor, with specimens of this species collected from around the world and with the holotypeof  D. hannibal, we could not find any significant morphological differences between these two species. As a result we consider  D. hannibalto be a junior synonym of  D. sulawensis.   Description.We provide a redescription to present a comparative uniform treatment of all species included in this paper.   Female: Holotypeof  Dyscritobaeus hannibalO’Connor et Ashe, Maputo,  V. 1989, leg. Angela( MSNG). Length of the body: 0.9 mm  Head. Color dark brown, lighter in the lower half. Coriaceous, frontal depression with a smooth area starting from the interantennal process and reaching the mid eye height. Centralkeel reaching 1/3 the height of the eye ( Fig. 73). Fan-like striation in malar area from oral foramen to eye margin, striation absent in genal area ( Fig. 74). Preoccipital lunula present and sculptured (Figs 3, 15, 75). Head width: height: length = 0.36: 0.31: 0.19 mm. Malar sulcus length: 0.1 mm. Eye width: height = 0.11: 0.16 mm. OOL: LOL: POL = 0.14: 0.05: 0.09 mm. Mandibles bidentate with dorsal tooth a little larger than ventral one ( Fig. 74). Palpi as in Fig IV 6, 7 after Caleca & Mineo (1995)( Fig. 44ain this paper). Antenna. Radicle and proximal half of A1 yellowish to light brown; distal half of A1 and proximal half of A2 brown; distal half of A2 light brown to yellow; A3 to A12 brown ( Fig. 73). A1 length: 0.19 mm; A2 longer as A3+ A4. Clava six-segmented; A11 is long, twice A12 length ( Fig. 17). Claval sensilla formula 0:2:2:2:1:0 (Fig. 18).  Mesosoma. In dorsal view, mesonotum and mesoscutellum dark brown with posterior mesoscutellar margin yellowish; propodeum dark red. In lateral view, pleurae dark red with metapleural carina yellowish ( Figs 75, 76). In dorsal view mesonotum and mesoscutellum imbricate; in lateral view speculum with furrow, femoral depression smooth, posterior mesepimeral area delimited anteriorly by a mesepimeral sulcus with a crenulate furrow, dorsal and ventral metapleural area with crenulate depression with rugae ( Fig. 76). Metapleural carina with two pointed projections, one adjacent to propodeal spiracle and second one at posterior margin ( Figs 75, 76). Metascutellum visible in dorsal view, triangular, large metanotal spines slightly surpassing propodeum ( Figs 75, 76). Mesosoma length 0.19 mm, width 0.35 mm; mesoscutellum length 0.11 mm, width 0.24 mm. Wings. Fore wing hyaline, surpassing metasoma; ratio st: pm: mg: sm: ww: lw: af: pf = 1: 1.3: 0.5: 3.4: 3.4: 8.3: 0.3: 1.0; angle st-pm= 23° as in Fig. 39a. st length is 0.08 mm. Coxae brown, femurs brown in the middle to yellowish in the tips; fore and middle tibiae brown in the middle to yellowish in the tips. Hind tibiae and tarsi yellow.  Metasoma. In dorsal view, tergites brown, in ventral view sternites brown, laterotergites yellowish. On T1 the pair of longitudinal submedial carinae are aligned with the metapleural carinae and separate lateral coriaceous microscupture from a microsculpure striate to coriaceous starting at basal grooves ( Fig. 11). T2 with foveolate anterior margin; costulate to coriaceous, rounded specillum in the middle of T2, costae converging to posterior margin of specillum ( Fig. 11).   Male(nov.): Singapore, Mandayroad,  1978/11/30, J. T. Huber, ( CNCI), [  CNC471528]. Length of the body: 0.71 mm  Head. Darker than in female, black ( Fig. 77). Sculpture as in female. Frontal depression shallower than in female; central keel little surpassing 1/3 the height of the eye. Fan-like striation in malar area from oral foramen to eye margin, striation just evident in genal area. Measures of the head width: height: length = 0.29: 0.25: 0.11 mm. Malar sulcus length: 0.07 mm. Measures of eye width: heigth = 0.09: 0.12 mm. OOL: LOL: POL = 0.06: 0.05: 0.05 mm. Mandibles as in female. Antenna. Radicles yellow, scape yellowish on tips, brown in the middle; A2– A12 brown; A12 two times A11. Sex segment with keel in the middle of antennomere ( Fig. 43)  Mesosoma. Color lighter than in female, with some parts brown yellow (Fig. 31). Sculpture as in female. Mesosoma length 0.19 mm, width 0.29 mm; mesoscutellum length 0.07 mm, width 0.19 mm. Wings. Fore wing hyaline, surpassing metasoma; ratio st: pm: mg: sm: ww: lw: af: pf = 1: 1.4: 0.4: 2.8: 3.1: 7.5: 0.4: 0.6; angle st-pm= 23° as in Fig. 39b. st length 0.11 mm  Metasoma. T1 and first half of T2 yellowish; last part of T2 and all the other tergites brown. Sculpture as in female except T2: specillum absent and costae slightly curving in the middle and converging at posterior margin of T2 (Figs 34, 78).   Comment and variability.This species has the widest distribution of all  Dyscritobaeusspecies. Four hundred and fifty-four specimens were examined from Africa, Indonesia, Japan, Malaysia, Nepal, Australia, Bangladesh, French Polynesia, India, Korea, New Caledonia, Papua New Guinea, Philippine, Singapore, Solomon Island, Sri Lanka. The degree of sexual dimorphism is also pronounced: specillum present in female and absent in male; female body dark brown, male with black head, mesosoma and metasoma with yellow parts in contrast with the rest brown or dark brown. Female length from 0.61 mmin Orientalspecimens to 1.19 mmin Japanese specimens. Females from the Australian region are a little smaller than African ones, the central keel is shorter, and the frontal depression a little bit shallower than in African specimens. Male body length from 0.64 mmin African specimens to 1.2 mmin Japanese specimens. Larger specimens of males are more sculptured than smaller ones. A2 completely yellow in smaller specimens, brown in the middle in larger specimens. Mesosoma in dorsal view may be brown. In some specimens the costae on T2 are curved in the middle of the posterior margin forming a small area resembling a faint specillum (Figs 34, 78). Anthony Watsham illustrated a dark  Dyscritobaeus(available in Johnson 2016:  Dyscritobaeus, image 306). This specimen is probably  D. sulawensis, considering that Watsham collected many  D. sulawensis(see material examined) and no other similar dark  Dyscritobaeusspecimens, but also in this case, as happened for  D. orientalisand probably for  D. indicus, he overlooked the presence of the specillum, illustrating T2 as being entirely striate (high magnification and correct lighting is critical to discern the specillum). The holotype of  D. hannibal,here considered to be a junior synonim of  D. sulawensis,is deposited in the collection of the Genoa Museum (MSNG) and not in the Insect Collection of Portici Università Federico II, Napoli, Italy (DEZA) as stated by O’Connor and Ashe (2011) in the original description.   Distribution.This species is recorded from the Oriental, Palearctic, Afrotropical, and Australian regions.    Material examined. Holotype. ♀:  INDONESIA:  Sulawesi: 0°34'N, 123°54'E,  14.x-1.xi.1985, Bogani Nani Wartabone National Park, over the first span of the footbridge over the  River Toraut, P.Ashe, Malaise Trap, ( NMID) [ NMINH:2016.7.1].   AFRICA:  Benin: Abomey-Calavica. 25km N. Coconou, 2♀, 1988/12, J. S. Noyes, MT, ( CNCI), [  CNC471382, 471383]; Abomey Calavi, ~25 Km NCotonou,  30♀Ƌ,  XII-1988 JS Noyes, MT, ( OSUC), [  OSUC148196, 148197, 148198, 418406, 418407, 418408, 418410, 418411, 418412, 418414, 418416, 418417, 418420, 418423, 418424, 418428, 418430, 418431, 418432, 418438, 418442, 418443, 418444, 418446, 418448, 418450, 418453, 418454, 418456, 418458];  Burkina Faso: Comoé,  Forêt de Boulon, 10°16'39''N 4°29'28''E,  270m, 10♀, 2006/07/07, F. & S. Génier, MT, (CNCI), [CNC 471384, 471385, 471386, 471387, 471388, 471389, 471390, 471391, 471392, 471393];   Cameroun: Nkoemvon, 1♀, 1979/07–1979/08, P. Jackson, MT, ( CNCI), [  CNC471395]; 1♀, 1979/09, D. Jackson, (BMNH), [BMNH (E)#971580]; 2♀, 1980/04, P. Jackson, MT, (CNCI), [CNC 471396, 471397]; 1♀, 1980/05/14– 1980/05/18, P. Jackson, MT, (CNCI), [CNC 471394];  Nkoemvon, Ebolowa, 2Ƌ, 1978/03–1978/05, D. Jackson, ( BMNH), [ BMNH(E)#971581, (E)#971582]; 3Ƌ, 1979/09, D. Jackson, ( BMNH), [ BMNH(E)#971583, (E)#971584, (E)#971585]; 1Ƌ, 1980/07–1980/08, Ms D.Jackson, ( BMNH), [ BMNH(E)#971586];  Central African Republic:  Parc National de Dzanga-Ndoki, 38.6km173º S Lidjombo, Lowland Rainforest, 2º21.60'N 16º09.20'E,  350m, 3♀, 5Ƌ, 2001/05/22, S. van Noort, Sweeping, (SAMC), [SAM- HYM-P 030679, P 030680, P025605, P025613, P025615, P025619, P047825, P047982]; 1Ƌ,   2001/05/23, S. van Noort, Sweeping, ( SAMC), [SAM-HYM-P025525];  Parc National de Dzanga-Ndoki, Mabéa Bai, 21.4 km53º NE Bayanga, Lowland Rainforest, marsh clearing, 3º02.01'N 16º24.57'E,  510m, 1♀, 2001/05/03, S. van Noort, Sweeping, (SAMC), [SAM-HYM-P025534]; 8Ƌ, 2001/05/05, S. van Noort, Sweeping, (SAMC), [SAM-HYM- P025552, P025558, P025559, P025561, P025565, P048101, P047977]; 2♀, 3Ƌ, 2001/05/06, S. van Noort, Sweeping, (SAMC), [SAM-HYM-P025536, P025546, P025566, P047830, P047979]; 1♀, 4Ƌ, 2001/05/06– 2001/ 05/06, S. van Noort, Sweeping, (SAMC), [SAM-HYM-P047824, P047980];  6♀, 5Ƌ,  2001/05/07, S. van Noort, Sweeping, ( SAMC), [SAM-HYM-P025575, P025576, P047826, P047827, P047828, P048100, P047978]; Réserve Spéciale de Forét Dense de Dzanga-Sangha, 12.7 kilometers 326°  North WestBayanga, Lowland Rainforest, 3° 00.27' N 16° 11.55' E,  420m, 1♀, 2Ƌ, 2001/05/12, S. van Noort, Sweeping, (SAMC), [SAM-HYM-P025589, P025594, P047829]; 1Ƌ, 2001/05/14– 2001/05/15, S. van Noort, Malaise Trap, (SAMC), [SAM-HYM-P025553]; 1Ƌ,   2001/05/17, S. van Noort, Sweeping, ( SAMC), [SAM-HYM-P047981]; Prefecture Sangha-Mbaéré P.Nat. de Dzanga-Ndoki 38.6km173°S Lidjombo, 2°21.60'N 16°03.20'E,  350m, 1♀Ƌ, 24-25.v.2001, S. van Noort Malaise rap, CAR01- M206 Lowland Rainforest, ( OSUC), [  OSUC344711];  Democratic Republic of the Congo:  Bandundu, Wamba, Kikongo Mission, along forest edge, 4°15'S 17°10'E, 1♀,  2006/04/19, S. L. Heydon, S. E. Stevenson, YPT, ( CNCI), [  CNC471398];  Yangambi, 1Ƌ,  1946/11/07, Catéier, ( IRSNB), [ I. G.24.778] [USNMENT01223068]; Yangambi, Desmodium Triflorum, Belvedere, 4Ƌ,   1946/11/07, ( IRSNB), [ I. G.24.778] [USNMENT01223069, 0 1223070, 0 1223071, 01223072];  Gabon: Réservedes Monts Doudou,  25.2 km304° NW Doussala, Forest-Coastallowland rainforest undergrowth - low canopy, 2° 13.63' S 10° 23.67' E,  660m,  1♀,  2000/ 03/15, S. van Noort, Sweeping, ( SAMC), [SAM-HYM-P048099];  Ghana: AshantiProv.,  Bobiri ForestReserve, 6°42'N 1°20'W, 1♀, 2001/07/23- 2001/07/31, C. Carlton, FIT, (CNCI), [CNC 471399]; 1♀, 2001/08/01–   2001/08/ 22, C. Carlton, FIT, ( CNCI), [  CNC471400]; 6°32'34"N 03°03'06"WBia National Park Camp 15, Researchcamp,  190m, 48♀Ƌ,   22-25.vi.2005 J.Gill& TK Phillipscoll., YPT, ( OSUC), [  OSUC495941, 495942, 495944, 495945, 495946, 495947, 495948, 495949, 495950, 495951, 495952, 495953, 495954, 495955, 495957, 495958, 495959, 495961, 495962, 495963, 495964, 495966, 495967, 495968, 495970, 495971, 495972, 495973, 495974, 495975, 495976, 495977, 495978, 495979, 495980, 495981, 495982, 495983, 495984, 495985, 495986, 495987, 495988, 495989, 495990, 495991, 495992, 495993];  Guinea: Mt-Nimba, rainforest, 7°41'N8°23W,  514m– 740m, 1♀, 1990/12–1991/03, L. Leblanc, FIT, (CNCI), [CNC 471401];  Mt-Nimba, Gouan River,  514m, 1♀, 1991/01/15–   1991/01/22, L. Leblanc, FIT, ( CNCI), [  CNC471402];  Ivory Coast: Bouake, 1♀, 1978/02/07, P. Cochereau, Malaise trap, (CNCI), [CNC 471403]; 2♀, 1978/06–1978/07, P. Cochereau, Malaise trap, (CNCI), [CNC 471409, 471410]; rice field, 4♀, 1980/02, P. Cochereau, (CNCI), [CNC 471411, 471412, 471413, 471414]; 1♀, 1980/02, P. Cochereau, mal. tr., (CNCI), [CNC 471415]; 3♀, 2Ƌ, 1980/02, P. Cochereau, pan trap, (CNCI), [CNC 471416, 471417, 471418, 471419, 471420]; 7♀, 1980/03, P. Cochereau, (CNCI), [CNC 471421, 471422, 471423, 471424, 471425, 471426, 471427]; rice field, 1♀, 1980/03, P. Cochereau, mal. tr., (CNCI), [CNC 471428]; 2♀, 1Ƌ, 1980/ 10/11, P. Cochereau, PT, (CNCI), [CNC 471404, 471405, 471406]; 2♀, 1981/10/01, P. Cochereau, PT, (CNCI), [CNC 471407, 471408];  1♀, 1981/12, P. Cochereau, PT, ( CNCI), [  CNC471429]; 6°13'N 5°02'WLamto ~ 160kmNW of Abidjan,  13♀Ƌ,  xi.1988 J.S.NoyesMT/YPT, ( OSUC), [  OSUC141958, 151718, 151719, 151721, 151722, 64644, 64687, 64743, 64782, 64804, 64820, 64829, 64857]; Lamto Fauchagesavane,  1♀Ƌ,  4.vi.1986 JY Rasplus, ( OSUC), [  OSUC56475];  Kenya:  Malindi, forest near coast, 1Ƌ, 1973/05, H. Falke, ( CNCI), [  CNC471430]; Nairobi,  1♀, 1Ƌ, 1982/02, M. C. Lubega, pan trap, ( CNCI), [  CNC471431, 471432(Ƌ, wing on slide,  Fig.39b)]; 2Ƌ, 1982/03, M. C. Lubega, pan trap, ( CNCI), [  CNC471433, 471434]; Nairobi, Ngong Rd., forest,  1♀,  1991/12/ 11, B. D. Gill, FIT, ( CNCI), [  CNC471435]; WesternProv.  Kakamega, 0°14.13'N 35°51.87'E, 1♀,  2000/06/19– 2000/06/26, R. Copland, MT, (CNCI), [CNC 471436]; ICIPE Mbita Point Feb.,  1♀Ƌ, 1983 M.C. Lubega, ( OSUC), [  OSUC56474]; Nairobi, KenyaInt. CenterIns. Physiol.&Ecol., 1♀ Ƌ, ii.1982MC Lubega, (OSUC), [OSUC 56470]; 2♀ Ƌ, iii.1982MC Lubega, (OSUC), [OSUC 56467, 56468]; 1♀ Ƌ, XI.1982MC Lubega, (OSUC), [OSUC 56473];   Mozambique: ( Holotype ♀+ 2 Paratypes ♀of  Dyscritobaeus hannibalO’Connor et Ashe) Maputo, 3♀, 1989/05, Angela, (MSNG) [USNMENT01223044, 0 1223045, 01223046];  2♀, 1993/05, R. Paci, ( UNIPA) [USNMENT01223047, 01223048]; NiassaCuamba, Mituque,  CentroPastoral, 1♀,  2012/03/28– 2012/04/3, M. Olmi, MT, (CNCI), [CNC 471437];   Nigeria: Ibadan, 1Ƌ,  1962/10/27, D. C. Eidt, ( CNCI), [  CNC471438]; 1♀, 1962/12/15, D. C. Eidt, (CNCI), [CNC 471439]; 1♀, 1962/12/20, D. C. Eidt, (CNCI), [CNC 471440]; 1Ƌ, 1962/12/22, D. C. Eidt, (CNCI), [CNC 471441]; 1Ƌ, 1962/12/26, D. C. Eidt, (CNCI), [CNC 471442]; 1♀, 1962/12/28, D. C. Eidt, (CNCI), [CNC 471443]; 1Ƌ,  1963/01/09, D. C. Eidt, ( CNCI), [  CNC471444]; OndoState, Idanre Hills, 7°06'16''N 05°06'13''E,  437m, 1♀, 2008/07/17, J. Hearty, sweeping, (CNCI), [CNC 471445];  OyoIbadan, IITAcompound,  1♀, 1987/11, J. S. Noyes, PT, ( CNCI), [  CNC471446]; KadunaSt. 81 kmNW Jos, 1♀ Ƌ, 11.xi.1987PNeuenschwander, (OSUC), [OSUC 56477];   Rwanda: Kibali, 1♀, 1977 ( UNIPA) [USNMENT01223049];   Seychelles: Denis I., 1Ƌ,  2000/04/13, M. J. Hill, MT, plot F1, ( CNCI), [  CNC471447];  Mahé, 4°37'S 50°26'E,  50m, 1♀,  2000/08/19, J. S. Noyes, ( CNCI), [  CNC471448];  Mahé, Tea Factory, 4°40'S  400m, 1♀,  2000/08/30, J. S. Noyes, ( CNCI), [  CNC471449];   South Africa: E. Transvaal, 11kmSE Pilgrims Rest, relict native forest,  1400m, 2♀,  1985/12/11–   1985/12/31, S. & J. Peck, FIT-Mal., ( CNCI), [  CNC471450, 471451]; Kruger Nat. Park, Pafuri, 12kmW jct. Levuvhu& LimpopoRivers, 2♀, 1984/04/01–   1984/04/21, M. Brigham, PT, ( CNCI), [  CNC471452, 471453]; Umtamvuna Nature Reserve, Forest - Coastal, 31° 03.509' S 30° 10.484' E,  160m, 1♀, 1Ƌ, 2000/11/11– 2000/11/18, S. van Noort, Pitfall Trap, (SAMC), [SAM-HYM-P 031743, P047984]; 2♀, 2000/11/11–   2000/11/18, S. van Noort, Yellow Pan Trap, ( SAMC), [SAM-HYM-P047832]; 28°23.1'S 32°24.4'E KwaZulu-Natal, St. Lucia, el.  10m, Gwalagwala Tr.  ANMTsite 1088, 11♀Ƌ,  7-20.ii.2004, Coastal Forestw/many lianas, A. Newton,  A. Solodovnikov & M. Thayer, flight intercept trap  FMHD#2004-044, ( OSUC), [  OSUC344663, 344664, 344665, 344666, 344667, 344668, 344669, 344670, 344671, 344672, 344673];   Tanzania: Dar es Salaam, Univ. Of Dar es Salaam, 1♀, 1984/10–1984/11, J. Middleton, ( CNCI), [  CNC471454]; E Usambara Mts., Amani NR, 05°10'34''S 38°36'01''E,  1004m, 1Ƌ,   2011/12/15, V. Grebennikov, sift. 0 5, ( CNCI), [  CNC471455]; IringaKilolo dist., Udzungwa Mts., Ndundulu Forest, Luwalacamp area, seasonal flooded meadow grassland surrounded by semi-evergreen montane virgin forest,  1880m, 1Ƌ, 2007/03/01–   2007/03/30, L. A. Hansen, MT, ( CNCI), [  CNC471456]; Mkomazi Game Reserve, Ibaya Camp, Acacia-Commiphora-Combretum bushveld, 3.58'S 37.48'E, 1Ƌ, 1996/01/29–   1996/03/11, S. van Noort, Malaise Trap, ( SAMC), [SAM-HYM- P018340]; Mkomazi Game Reserve, Kinondo Mtn, Forestedge - Montane,  1♀,  1996/05/09, H.G. Robertson, Winkler Bag, ( SAMC), [SAM-HYM-P018847]; Udzungwa Mts., -7,8447 36,8677,  1113m, 1Ƌ, 2014/10/07, V. Grebennikov, sift. 0 2, (CNCI), [CNC 471457];  Zanzibar, Prison Island, Changu, 1♀, 1995/05, Ann Helen Renning, ( CNCI), [  CNC471458];   Uganda: 13kmSE Kampala, Gabawetland, 1♀,  2004/06/10–   2004/07/17, I. Přikryl, Malaise Trap, ( CNCI), [  CNC471459]; KibaleNational Park, Kanyawara, Makerere, University Biological Field Station, Primarymid-altitude Rainforest, near stream,  1587m, 1♀, 2008/08/04–   2008/08/26, S. van Noort, Yellow Pan Trap, ( SAMC), [SAM-HYM-P  031257]; Secondarymid-altitude Rainforest, 0° 33.871' N 30° 21.355' E,  1495m, 2♀, 2008/08/04–   2008/08/26, S. van Noort, Yellow Pan Trap, ( SAMC), [SAM-HYM-P047831]; Secondarymid-altitude Rainforest, marshy area, 0° 34.806' S 30° 21.874' E,  1491m, 1♀, 1Ƌ, 2008/08/02–   2008/08/12, S. van Noort, Malaise Trap, ( SAMC), [SAM-HYM-P  030696]; Primarymid-altitude Rainforest, near stream, 0°34.405'N 30°21.646'E,  1484m, 2Ƌ, 2008/08/04–   2008/08/26, S. van Noort, Yellow Pan Trap, ( SAMC), [SAM-HYM- P047983];  Yemen: Ar Rujum, 1♀, 2000/10/16–   2001/01/15, A. van Harten, MT, ( CNCI), [  CNC471460]; Khamis Bani Sa'd, litter, 1Ƌ,   1999/06/09, A. van Harten, ( CNCI), [  CNC3754/471461]; Lahj, 1Ƌ, 2000/09,  A. van Harten, ( CNCI), [  CNC5076/471462];   Zimbabwe: Harare(Salisbury), 2♀, 1980/04,  A. Watsham, pantr., ( CNCI), [  CNC471495, 471496];  4♀, 1Ƌ, 1980/04, A. Watsham, PT, ( CNCI), [  CNC471463, 471464, 471465, 471466, 471467];  Zimbabwe(Rhodesia): 1Ƌ, 1975/05–1975/07,  A. Watsham, ( CNCI), [  CNC471468];  1♀, 1979/04, A. Watsham, ( BMNH), [ BMNH(E)#971587];  3♀, 4Ƌ, 1979/05, A. Watsham, ( CNCI), [  CNC471469, 471470, 471471, 471472, 471473, 471478, 471479];  3♀, 2Ƌ, 1979/06, A. Watsham, ( CNCI), [  CNC471476, 471477, (E)#971588, 471474, 471475];  1♀, 1979/07, A. Watsham, ( CNCI), [  CNC471480];  1♀, 1980/02, A. Watsham, Pan Trap, ( CNCI), [  CNC471497];  2♀, 1980/03, A. Watsham, Pan Trap, ( CNCI), [  CNC471498(wing on slide, Fig. 39a), 471499];  14♀, 1980/05, A. Watsham, ( CNCI), [  CNC471484, 471485, 471486, 471487, 471488, 471489, 471490, 471491, 471492, 471493, 471494, 471481, 471482, 471483];  3♀, 1980/06, A. Watsham, ( CNCI), [  CNC471500, 471501, 471502];  2♀, 1980/07, A. Watsham, ( CNCI), [  CNC471503, 471504];  Harare(Salisbury), Chishawasha, 1♀, 1979/ 10,  A. Watsham, ( CNCI), [  CNC471505]; 8♀, 1Ƌ, 1980/02,  A. Watsham, pan trap, ( CNCI), [  CNC471506, 471507, 471508, 471509, 471510, 471511, 471512, 471513, 471514];  10♀, 3Ƌ, 1980/03, A. Watsham, ( CNCI), [  CNC471515, 471516, 471517, 471518, 471519, 471520, 471521, 471522, 471523, 471524, 471525, 471526, 471527];   AUSTRALIA:  Queensland: 3kmNE Mt.  Webb, 2♀,  1980/10/01–   1980/10/03, J.C. Cardale, Yellow Pan Trap, ( ANIC), [  ANIC32- 070476, 32- 070477]; s. s. Rain forestundergrowth, 1♀, 1981/04/30– 1981/05/04, J. C. Calder & J. Feehan, Berlesate Trap, (ANIC), [ANIC 32- 070478];  Base Tinaroo Falls, 1Ƌ,  1976/03/30, I. D. Galloway, ( QDPC) [USNMENT01223050]; Bellenden Ker Range, Cableway Base Stn,  100m, 2♀, 1981/10/17–   1981/11/09, Baited window Trap, ( QDPC) [USNMENT01223051, 01223052]; Bowenia State Forest, Byfield Cent. Q., 1♀, 1Ƌ, 1975/10/31, I. D. Galloway, D-vac., (QDPC) [USNMENT01223053, 01223054]; Brookfield Nr Brisbane, 1♀, 1982/01, Boucek, (BMNH), [BMNH (E)#971589]; Cairns, 1Ƌ, (ANIC), [ANIC 32- 070479]; Capalaba, 2Ƌ, 1980/06/08, J. S. Noyes, (CNCI), [CNC 471590, 471591]; Daintree Nat. Park, CapeTribulation, Bloomfield Trackat Emmagen creek, rainforest, 16°020.357'S 145°27.443'E,  30m, 1♀,  2005/12/17–   2005/12/18, A. R. Deans & M. Buffington, yellow, blue, red pan traps(4:1:1), ( CNCI), [  CNC471592]; Eurimbula Ck., via Round Hill Head, 1♀, 1975/05/03–   1975/05/05, I. Naumann, ( QDPC) [USNMENT01223055]; Gap. Ck. 5kmESE  Mt. FinniganQ., 3♀, 1981/05/13–   1981/05/16, I. D. Naumann, Yellow Pan Trap, ( ANIC), [  ANIC32- 070480, 32- 0 70481, 32-  070482]; Gatton, 1♀, 1Ƌ, 1981/03/30– 1981/04/06, Yellow Pan Trap, (QDPC) [USNMENT01223056, 01223057]; 1♀, 1981/04/06, D-vac in potato crop, (QDPC) [USNMENT01223058]; Grap Ck. 5kmESE, Mt. Finnigan, 6Ƌ, 1981/05/13– 1981/05/16, I. D. Naumann, Yellow Pan Trap, (ANIC), [ANIC 32- 070483, 32- 070484, 32- 070485, 32- 070486, 32- 070487, 32- 070488]; Great Sandy Nat. Park, 25°00.62'S 153°02.80'E, 1Ƌ, 2002/12/16–   2002/12/17, J. Munro, A. Owen, YPT, ( CNCI), [  CNC471593];  Laceys CreekQMission Beach, s. s. Rain forestundergrowth, 1♀, 1Ƌ, 1980/05/13–   1980/05/14, I. D. Naumann& J. C. Cardale, ( ANIC), [  ANIC32- 070489, 32- 070490]; Mossman Gourge,  30m, 2♀, 1984/02/23, L. Masner, (CNCI), [CNC 476269, 476270]; Mt Webb Nat. Pk., 2Ƌ, 1980/09/28– 1980/09/30, T. Weir, Berlesate Trap, (ANIC), [ANIC 32- 070491, 32- 070492]; 1Ƌ, 1981/04/27– 1981/04/30, I. D. Naumann, Yellow Pan Trap, (ANIC), [ANIC 32- 070493];  Mt. Tamborine, 1♀, 1978/11–1979/01, Pitfall Trap, ( QDPC) [USNMENT01223059];  Mt. Webb, 4♀,  1981/04/27– 1981/04/30, I. D. Naumann, Yellow Pan Trap, (ANIC), [ANIC 32- 070494, 32- 070495, 32- 070496, 32- 070497];  Pebbly Beach, Nof Cairns, 1Ƌ,  1974/04/ 27, J. F. Donaldson, ( QDPC) [USNMENT01223060]; Rotary Park, Mossman, 1♀, 1975/11/06, I. D. Galloway, Dvac., (QDPC) [USNMENT01223061]; Samford, 1Ƌ, 1962/01, E. Warwiell, Yellow Pan Trap, (ANIC), [ANIC 32- 070498];  Shiptons Flat, 1♀,  1980/10/17–   1980/10/19, J.C. Cardale, ( ANIC), [  ANIC32- 070499]; Townsville James Cook Uni Campus, 1Ƌ, 1988/05/21–   1988/05/24, A. D. Austin, Pitfall Trap, ( WINC) [USNMENT01223062];   Tully FallsRoad, 1♀,  1976/03/31, I. D. Galloway, ( QDPC) [USNMENT01223063];    WesternAustralia: “ Marun” CALMSite 8/4 Prince Frederick Harbour, 1Ƌ,  1988/06/06–   1988/06/11, I. D. Naumann, Yellow Pan Trap, ( ANIC), [  ANIC32- 070500]; Mining Camp, Mitchell Plateau, 3♀, 1Ƌ, 1983/05/09–   1983/05/19, I. D. Naumann& J. C. Cardale, Yellow Pan Trap, ( ANIC), [  ANIC32- 070501, 32- 070502, 32-  070503, 32-  070504];  BANGLADESH:   Bangladesh: potato crop, 1♀,  1982/11/03, J. V. Harten, PT, ( CNCI), [  CNC471531]; FIJI: Suva: Nabua,  SPC Compound, 1♀,  2002/06/22–   2002/06/30, L. Leblanc, ( CNCI), [  CNC471532];  FRENCH POLYNESIA:  Bora Bora: nr. Pt. Paoaoa, 1♀, 2Ƌ,   1984/09/04, D. M. Lasalle, ( CNCI), [  CNC471533, 471534, 471535];  Tahitii: Bora Bora, in grass, 3♀,   1982/07/21, H. Andersen Col., sweeping grasses, ( CNCI), [  CNC471536, 471537, 471530];  INDIA:   Karnataka: Bangalore, CIBCgrounds, 1♀, 1974/12/16, PT, (CNCI), [CNC 471529]; Uttar Predesh: Aligarh, 1♀, 1980, M. Hayat, (CNCI), [CNC 471538]; 1♀, 1980/11, M. Hayat, (CNCI), [CNC 471539];  1♀, 1980/ 12, M. Hayat, ( CNCI), [  CNC471540];  INDONESIA:   Bali: Penulisan, potato crop, 1♀,  1978/11/21, J. T. Huber, ( CNCI), [  CNC471541];  Curug: 1♀, 1980/03/14– 1980/03/21, H. R. Gillespie, pan trap, (CNCI), [CNC 471542]; 2♀, 1980/12/20–   1980/12/30, H. R. Gillespie, pan trap, ( CNCI), [  CNC471543, 471544];  Papua: Neth. Guega, W.of Swart Val.,  1200m, 1Ƌ,   1958/11/14, J. L. Gressitt, ( BPBM) [USNMENT01223064];  Sulawesi Utara: Dumoga Bone Nat. Pk., Sievedrain-forest litter,  220m, 2♀, 1985/06/07–   1985/06/25, A. D. Austin, ( WINC) [USNMENT01223065, 01223066];   JAPAN:  Fukuoka: Mt. Tachibanayama, 2Ƌ,  1980/08/12, C. M. Yoshimoto, ( CNCI), [  CNC471545, 471546];   GunmaPref.: Takamine Table Land,  1950m, 1♀,   1980/07/19, A. & Z. Smetana, ( CNCI), [  CNC471376]; Honshu Aichi: Yakusa, Toyota, 2♀, 1990/07/30–   1990/08/07, K. Yamagishi, PT, ( CNCI), [  CNC471272, 471273]; Honshu IbarakiPref.: Tsuchiura, 1Ƌ, 1989/08/08– 1989/08/13, M. J. Sharkey, (CNCI), [CNC 471274];  Tsuchiura forest, 1♀,  1989/09/11– 1989/09/19, M. J. Sharkey, PT, (CNCI), [CNC 471275]; 3♀, 1989/09/19– 1989/10/02, M. J. Sharkey, PT, (CNCI), [CNC 471276, 471277, 471278]; 1♀, 1989/10/02– 1989/10/ 16, M. J. Sharkey, PT, (CNCI), [CNC 471279]; 1♀, 1989/10/16–   1989/10/27, M. J. Sharkey, PT, ( CNCI), [  CNC471280]; Tsuchiuramarsh, 1Ƌ, 1989/09/27–   1989/10/03, M. J. Sharkey, PT, ( CNCI), [  CNC471281]; Tsuchiura, Shizizuka forest, 2♀, 1996/09/04–   1996/09/05, L. Masner, YPT, ( CNCI), [  CNC471282, 471283]; Tsukuba, 6♀, 1989/05/26–   1989/05/31, M. Sharkey, YPT&FIT, ( CNCI), [  CNC471284, 471285, 471286, 471287, 471288, 471289]; Tsukuba, Matsushiro, 1Ƌ, 1989/07/26–   1989/08/04, M. J. Sharkey, PT, ( CNCI), [  CNC471290]; Honshu TochigiPref.: Nishinasuno, N.G.R.I.,  500m, 3♀, 1Ƌ,   1989/08/10, M. J. Sharkey, sweep, ( CNCI), [  CNC471291, 471292, 471293, 471294];  Ibaraki: Mt. Tsukuba, 1♀, 1989/08/29–   1989/09/06, M. J. Sharkey, ( CNCI), [  CNC471295]; Tsuchiura City, Shishizuka-Oika, 1♀, 1989/08/13–   1989/08/25, M. J. Sharkey, MT, ( CNCI), [  CNC471296]; Tsuchiuramarsh, 1♀, 1989/06/24– 1989/06/31, M. J. Sharkey, Pan Trap, (CNCI), [CNC 471297]; 1♀, 1989/08/21– 1989/08/29, M. J. Sharkey, PT, (CNCI), [CNC 471298]; 1♀, 1Ƌ, 1989/08/29– 1989/09/06, M. J. Sharkey, PT, (CNCI), [CNC 471299, 471300]; 1♀, 1989/09/19–   1989/10/02, M. Sharkey, PT-FIT, ( CNCI), [  CNC471301]; Tsuchiura, Expo Site, 4♀, 8Ƌ, 1989/08/01–   1989/08/08, M. J. Sharkey, PT, ( CNCI), [  CNC471302, 471303, 471304, 471305, 471306, 471307, 471308, 471309, 471310, 471311, 471312, 471313]; Tsukuba, Expo Site, 5♀, 1989/07/24– 1989/07/31, M. Sharkey, PT, (CNCI), [CNC 471314, 471315, 471316, 471317, 471318]; 6♀, 2Ƌ, 1989/08/08– 1989/08/15, M. Sharkey, PT, (CNCI), [CNC 471319, 471320, 471321, 471322, 471323, 471324, 471325, 471326]; 8♀, 1989/08/15– 1989/08/29, M. Sharkey, PT, (CNCI), [CNC 471327, 471328, 471329, 471330, 471331, 471332, 471333, 471334]; 9♀, 2Ƌ, 1989/08/29– 1989/09/05, M. Sharkey, PT, (CNCI), [CNC 471335, 471336, 471337, 471338, 471339, 471340, 471341, 471342, 471343, 471344, 471345]; 1Ƌ, 1989/09/05– 1989/09/ 11, M. Sharkey, MT, (CNCI), [CNC 471346]; 1♀, 1Ƌ, 1989/09/05– 1989/09/11, M. Sharkey, PT, (CNCI), [CNC 471347, 471348]; 1Ƌ, 1989/09/18– 1989/09/24, M. Sharkey, PT, (CNCI), [CNC 471349]; 1♀, 1989/09/24– 1989/ 10/02, M. Sharkey, PT, (CNCI), [CNC 471350]; 1♀, 1989/10/27–   1989/11/03, M. Sharkey, PT, ( CNCI), [  CNC471351]; Tsukuba, NIAES, 2Ƌ, 1989/06/26– 1989/07/10, M. Sharkey, PT, (CNCI), [CNC 471352, 471353]; 1♀, 1989/07/10– 1989/07/17, M. Sharkey, PT, (CNCI), [CNC 471354]; 6♀, 2Ƌ, 1989/07/14– 1989/07/25, M. Sharkey, PT, (CNCI), [CNC 471355, 471356, 471357, 471358, 471359, 471360, 471361, 471362]; 2♀, 2Ƌ, 1989/08/04– 1989/08/14, M. Sharkey, PT, (CNCI), [CNC 471363, 471364, 471365, 471366]; 1♀, 1989/08/14– 1989/08/27, M. Sharkey, PT, (CNCI), [CNC 471367]; 5♀, 1Ƌ, 1989/09/07– 1989/09/11, M. Sharkey, PT, (CNCI), [CNC 471368, 471369, 471370, 471371, 471372, 471373]; 1♀, 1989/09/11–   1989/09/18, M. Sharkey, PT, ( CNCI), [  CNC471374];  Kyushu: Fukuoka, Mt Hiko, potato crop, 4♀, 1989/05/09– 1989/05/10, M. J. Sharkey, sweep, (CNCI), [CNC 471547, 471548, 471549, 471550]; 700m, 1♀, 1989/08/25– 1989/09/04, K. Takeno, M. Sharkey, MT, (CNCI), [CNC 471551]; 2♀, 1989/09/18– 1989/09/25, K. Takeno, M. Sharkey, MT, (CNCI), [CNC 471552, 471553]; 1♀, 1989/09/25–   1989/10/02, K. Takeno, M. Sharkey, MT, ( CNCI), [  CNC471554]; Fukuoka, Mt. Tachibana, 1♀, 1979/ 07/15– 1979/07/21, K. Yamagishi, cutover land, (CNCI), [CNC 471555]; 3♀, 1Ƌ, 1979/08/12– 1979/08/18, K. Yamagishi, YPT, (CNCI), [CNC 471556, 471557, 471558, 471559]; 2♀, 1Ƌ, 1979/08/19– 1979/08/25, K. Yamagishi, YPT, (CNCI), [CNC 471560, 471561, 471562];  4♀,  1992/08/17, K. Yamagishi, ( CNCI), [  CNC471563, 471564, 471565, 471566];  NaganoPref.: Nakakaruizawa,  1000m, 1Ƌ,   1980/07/19, A. & Z. Smetana, ( CNCI), [  CNC471375];  NEPAL:  Kakani:  2700m, 1♀, 1983/08/01– 1983/08/23, H. J. Allen, Malaise Trap, (CNCI), [CNC 471567];   NEW CALEDONIA:  Noumea: Ouen Toro, 2Ƌ, 1972/01, P. Cochereau, ( CNCI), [  CNC471571, 471572]; 1♀, 1Ƌ, 1972/01/07, P. Cochereau, (CNCI), [CNC 471568, 471569]; 1Ƌ, 1972/05, P. Cochereau, (CNCI), [CNC 471573]; 1♀, 1972/06/01– 1972/06/19, P. Cochereau, (CNCI), [CNC 471570]; PAPUA NEW GUINEA:   Madang: Morox, 4°16'S 144°58'E,  100m, 1♀,  2006/08/01–   2006/08/16, V. Iwam, YPT, ( CNCI), [  CNC471574]; Nagada, 5°17'S 145°49'E,  5m, 1Ƌ,   2006/06/06, J. Hrček, YPT, ( CNCI), [  CNC471575]; Wanang, 5°15'S 145°16'E,  100m, 2Ƌ,   2006/08/23, P. Damen, MT, ( CNCI), [  CNC471576, 471577];  MorobeProv: Wau Ecol. Instit, 2♀, 1983/07/26–   1983/07/31, S. & P. Miller, PT, ( CNCI), [  CNC471578, 471579]; 1Ƌ, 1983/08, S. & P. Miller, PT, ( CNCI), [  CNC471580];  Okapa: EasternH’ land,  South, 1Ƌ,  1964/10/27, R. Hornabrook, ( ANIC), [  ANIC32- 070505];   PHILIPPINES:  Davao:  Davao, 1♀, 1961/11, M. Gavarra, Yellow Pan Trap, ( BMNH), [ BMNH(E)#971590];   Negros Oriental:  Cuernos de Negros,  7kmW Valencia, primany forest edge, 9°17'N 123°15'E,  700m, 1♀,  1987/07/15– 1987/07/21, D. C. Darling, E. Mayordo, MT with pans, (CNCI), [CNC 471581]; 3♀, 1987/07/25–   1987/07/29, D. C. Darling, E. Mayordo, MT wit pans, ( CNCI), [  CNC471582, 471583, 471584];  SINGAPORE: Singapore: Manday road, 1Ƌ,   1978/11/30, J. T. Huber, ( CNCI), [  CNC471528]; SOLOMON ISLAND:  Guadalcanal: Honiara,   0–100m, 1Ƌ, 1970/10, N. L. H. Krauss, ( BPBM) [USNMENT01223067]; New Georgia: Munda, 1Ƌ, 1980/11, N. L. H. Krauss, ( CNCI), [  CNC471585];   SOUTH KOREA:  Chung-nam: Daejon-si, Wadong, for. Edge, wild rose patch, 36°24.02'N 127°25.98'E, 1♀,  2006/05/06–   2006/05/28, P. Tripotin, MT, ( CNCI), [  CNC471377];  Jirisan: Hamyang-gum, Macheon-myon, Samjeong-li, forest gully, 35°20'55''N 127°38'21''E,  700m, 2♀, 2002/08/25–   2002/09/14, P. Tripotin, MT, ( CNCI), [  CNC471378, 471379];  Kangwon: Chuncheon, Nam-myeon, Hudong-li, pastured area, trail close to forest edge, 1♀, 2003/06/14– 2003/07/06, P. Tripotin, MT, (CNCI), [CNC 471380]; in semi-shade, forest edge, 1♀, 2003/07/31–   2003/08/16, P. Tripotin, MT, ( CNCI), [  CNC471381]; district On-dzong, Kus-gangsan, along Ok-rudong,  300–600m, 1Ƌ,   1975/08/05, J. Papp& A. Vojnits, ( HNHM), [ No.317] [USNMENT01223073]; SRI LANKA:   Sabaragamuwa: Kegalle, 1♀,  1970/01/ 14, Mussard Besuchet Lobl, ( CNCI), [  CNC471588];   WestProvince: Nugegoda, 1Ƌ,  1969/01/01, B. P. Karumaratne, ( CNCI), [  CNC471589];  THAILAND:  PhetchabunKhao: Kho Nat. Park, 16°32.546'N0', 560m, 1♀, 2007/06/19–   2007/06/26, Somchai Chachumnan, S.Singtong, ( CNCI), [  CNC#2458/471586];  Suphanburi: Pu Toei Nat. Park, Huai Mongpae, 14°56.574'N 99°25.23'E, 378m, 1♀, 2008/09/08– 2008/09/09, W. Jaisue, PT, (CNCI), [CNC #3570/471587]. 1323915675 [199,1377,1722,1747] 1989-05 MSNG Angela Female 28 29 1 Maputo holotype 1323915693 [199,1186,908,933] 1978-11-30 CNCI, CNC J. T. Huber Singapore Male Manday 29 30 CNC 471528 1 1323915689 1985-10-14 1985-11-01 1985-10-14 NMID, NMINH Bogani Nani Wartabone National Park & River Toraut, P. Indonesia 0.56666666 River Toraut 1308 123.9 31 32 1 1 Sulawesi holotype 1323915691 CNCI, CNC Abomey-Calavi & N. Coconou & J. S. Noyes & Km N Benin Benin 31 32 CNC 471382, 471383 2 2 1323915687 1988-12 OSUC JS Noyes, MT & Foret de Boulon Burkina Faso 270 10.2775 Foret de Boulon 21 4.491111 31 32 OSUC 148196, 148197 30 30 Comoe 1323915709 CNCI, CNC Nkoemvon & P. Jackson Cameroon Cameroun 31 32 CNC 471395 1 1 1323915715 BMNH Nkoemvon & Ebolowa & D. Jackson & Ms D. & Parc National de Dzanga-Ndoki & S Lidjombo Central African Republic 350 2.3600001 Lowland Rainforest 13 16.153334 Parc National de Dzanga-Ndoki 31 32 1 1323915694 2001-05-23 SAMC S. van Noort & Sweeping & Parc National de Dzanga-Ndoki & NE Bayanga Central African Republic 510 3.0335 Lowland Rainforest 13 16.4095 Parc National de Dzanga-Ndoki 31 32 1 Bai 1323915680 2001-07-05 SAMC S. van Noort & Sweeping & Reserve Speciale de Foret Dense de Dzanga-Sangha & Lowland Rainforest Central African Republic 420 32 33 3.0045 North West 13 16.1925 31 32 6 6 North West 1323915695 2001-05-17 SAMC S. van Noort & Sweeping & P. Nat. de Dzanga-Ndoki Central African Republic 350 2.3600001 Lidjombo 13 16.053333 32 33 1 1 Sangha-Mbaere 1323915716 [303,1428,332,357] OSUC Lowland Rainforest Democratic Republic of the Congo Democratic Republic of the Congo 32 33 OSUC 344711 1 1323915696 2006-04-19 CNCI, CNC Wamba & Kikongo Mission & S. L. Heydon & S. E. Stevenson Democratic Republic of the Congo -4.25 Bandundu 1306 17.166666 32 33 CNC 471398 1 1 Bandundu 1323915700 1946-07-11 IRSNB Yangambi & Cateier & I.D. Galloway Democratic Republic of the Congo Yangambi Belvedere Desmodium Triflorum 32 33 1 Bandundu 1323915677 1946-07-11 IRSNB I.D. Galloway Gabon Reserve 660 -2.2271667 Forest-Coastal 13 10.3945 Monts Doudou 32 33 1 1323915703 2000-03-15 SAMC S. van Noort & Sweeping & Bobiri Forest Reserve Ghana 6.7 Bobiri Forest 1304 -1.3333334 32 33 1 1 Ashanti 1323915699 2001-08-22 CNCI, CNC C. Carlton Ghana 190 6.5427775 Research 21 -3.0516667 32 33 CNC 471400 48 48 Ashanti 1323915710 2005-06-22 2005-06-25 2005-06-22 OSUC J. Gill & TK Phillips Guinea-Bissau 514 7.6833334 Mt-Nimba 32 33 OSUC 495941, 495942 1 1323915688 [860,1227,836,861] Guinea-Bissau 514 Mt-Nimba 32 33 1 River 1323915698 [151,1058,871,896] 1991-01-22 CNCI, CNC L. Leblanc Ivory Coast Bouake 32 33 CNC 471402 1 1323915707 CNCI, CNC P. Cochereau Ivory Coast 6.2166667 Abidjan 1305 -5.0333333 32 33 CNC 471429 1 1 Abidjan 1323915708 1988-11 OSUC J. S. Noyes Ivory Coast Lamto Fauchage 32 33 OSUC 141958, 151718 13 13 Abidjan 1323915704 1986-06-04 OSUC JY Rasplus Kenya Kenya 32 33 OSUC 56475 1 1 1323915701 CNCI, CNC H. Falke Kenya Malindi 32 33 CNC 471430 1 1323915686 CNCI, CNC M. C. Lubega & Fig. Kenya Ngong Rd. 32 33 CNC 471431, 471432, CNC 471433, 471434 1 1 1323915702 1991-11-12 CNCI, CNC B. D. Gill Kenya Western 32 33 CNC 471435 1 1 Western 1323915697 [842,1422,1340,1365] 2000-06-19 Kenya 0.23550001 Kakamega 13 35.8645 32 33 1 1 Kakamega 1323915720 OSUC M. C. Lubega Kenya Center 32 33 OSUC 56474 1 1 Center 1323915714 Mozambique Maputo 32 33 4 2 Maputo holotype 1323915717 UNIPA R. Paci Mozambique Mituque 32 33 2 2 Niassa holotype 1323915713 [1049,1422,1556,1581] 2012-03-28 Mozambique Centro 32 33 1 1 Centro holotype 1323915734 1962-10-27 CNCI, CNC Ibadan & D. C. Eidt Nigeria Nigeria 32 33 CNC 471438 1 holotype 1323915741 1963-09-01 CNCI, CNC D. C. Eidt Nigeria 437 7.1044445 Idanre Hills 21 5.103611 32 33 CNC 471444 1 Ondo holotype 1323915722 [335,667,1772,1797] IITA Nigeria Oyo 32 33 1 Oyo holotype 1323915706 CNCI, CNC J. S. Noyes Nigeria Jos 32 33 CNC 471446 1 1 Kaduna holotype 1323915718 UNIPA Kibali Rwanda Rwanda 32 33 1 1 holotype 1323915740 2000-04-13 CNCI, CNC Denis I. & M. J. Hill Seychelles Seychelles 32 33 CNC 471447 1 holotype 1323915725 [260,1205,1879,1905] 2000-08-19 CNCI, CNC J. S. Noyes India 50 -4.616667 Mahe 1306 50.433334 32 33 CNC 471448 1 1 holotype 1323915719 2000-08-30 CNCI, CNC Tea Factory & J. S. Noyes India 400 -4.6666665 Mahe 32 33 CNC 471449 1 1 holotype 1323915705 1985-11-12 E. Transvaal South Africa 1400 Pilgrims Rest 32 33 2 2 holotype 1323915729 1985-12-31 CNCI, CNC S. & J. Peck South Africa Levuvhu Pafuri 32 33 CNC 471450, 471451 1 Limpopo holotype 1323915721 1984-04-21 CNCI, CNC M. Brigham South Africa 160 33 34 -31.058483 Forest - Coastal 1 30.174734 Umtamvuna Nature Reserve 32 33 CNC 471452, 471453 1 Limpopo holotype 1323915736 2000-11-18 SAMC S. van Noort Saint Lucia 10 -28.385 KwaZulu-Natal 123 32.406666 33 34 1 KwaZulu-Natal holotype 1323915712 2004-02-07 2004-02-20 2004-02-07 ANMT Saint Lucia Coastal Forest 33 34 11 11 A holotype 1323915723 FMHD, OSUC M. Thayer Saint Lucia A 33 34 FMHD#2004-044, OSUC 344663, 344664 1 A holotype 1323915732 CNCI, CNC Univ. Of & J. Middleton & E Usambara Mts. & Amani NR Tanzania 1004 -5.1761107 Usambara Mts. 21 38.600277 33 34 CNC 471454 1 1 Dar es Salaam holotype 1323915731 2011-12-15 CNCI, CNC V. Grebennikov Tanzania Udzungwa Mts. 1880 Luwala Ndundulu Forest 33 34 CNC 471455 1 Iringa holotype 1323915733 2007-03-30 CNCI, CNC Hansen, MT Tanzania Ibaya Camp Mkomazi Game Reserve 33 34 CNC 471456 1 A holotype 1323915726 1996-11-03 SAMC S. van Noort Tanzania Mkomazi Game Reserve Forest Kinondo Mtn 33 34 1 A holotype 1323915711 1996-09-05 SAMC H. G. Robertson & Winkler Bag Tanzania 1113 Udzungwa Mts. 33 34 1 1 A holotype 1323915728 CNCI, CNC Changu & Ann Helen Renning Tanzania Prison Island 33 34 CNC 471458 1 1 holotype 1323915724 [447,1164,692,717] 2004-10-06 Uganda Gaba 33 34 1 1 Kampala holotype 1323915744 2004-07-17 CNCI, CNC I. Prikryl Uganda Kanyawara 1587 University Biological Field Station Makerere 33 34 CNC 471459 1 Kibale holotype 1323915738 2008-08-26 57-03-12 2008-08-26 SAMC S. van Noort Uganda 1495 0.56451666 Rainforest 1 30.355917 Secondary 33 34 1 Kibale holotype 1323915742 2008-08-26 SAMC S. van Noort Uganda 1491 -0.5801 Rainforest 1 30.364567 Secondary 33 34 1 Kibale holotype 1323915737 2008-12-08 96-03-06 2008-12-08 SAMC S. van Noort Uganda 1484 0.57341665 Rainforest 1 30.360767 Primary 33 34 1 Kibale holotype 1323915759 2008-08-26 SAMC S. van Noort Uganda Ar Rujum Yem 33 34 1 Kibale holotype 1323915763 2001-01-15 CNCI, CNC van Harten, MT Uganda Khamis Bani Sa'd 33 34 CNC 471460 1 A holotype 1323915747 [388,1259,1015,1040] 1999-09-06 CNCI, CNC Uganda Lahj 33 34 CNC 3754 1 A holotype 1323915730 CNCI, CNC Uganda A 33 34 CNC 5076 1 A holotype 1323915748 [502,1019,1052,1077] Zimbabwe Harare 33 34 2 2 Harare holotype 1323915767 CNCI, CNC Zimbabwe A 33 34 CNC 471495, 471496 1 A holotype 1323915746 CNCI, CNC Watsham, PT Zimbabwe A 33 34 CNC 471463, 471464 4 4 A holotype 1323915743 [690,1119,1124,1149] CNCI, CNC Zimbabwe A 33 34 CNC 471468 1 A holotype 1323915727 BMNH Zimbabwe A 33 34 1 1 A holotype 1323915754 CNCI, CNC Zimbabwe A 33 34 CNC 471469, 471470 3 3 A holotype 1323915750 CNCI, CNC Zimbabwe A 33 34 CNC 471476, 471477 3 3 A holotype 1323915764 [259,836,1231,1256] CNCI, CNC Zimbabwe A 33 34 CNC 471480 1 1 A holotype 1323915739 CNCI, CNC Zimbabwe A 33 34 CNC 471497 1 1 A holotype 1323915745 [261,1366,1268,1293] CNCI, CNC Zimbabwe A 33 34 CNC 471498 2 2 A holotype 1323915757 CNCI, CNC Zimbabwe A 33 34 CNC 471484, 471485 14 14 A holotype 1323915756 CNCI, CNC Zimbabwe A 33 34 CNC 471500, 471501 3 3 A holotype 1323915758 [259,930,1375,1400] CNCI, CNC Zimbabwe A 33 34 CNC 471503, 471504 2 2 A holotype 1323915752 Zimbabwe Harare 33 34 1 1 Harare holotype 1323915735 [198,864,1412,1437] CNCI, CNC Zimbabwe A 33 34 CNC 471505 8 8 A holotype 1323915751 CNCI, CNC Zimbabwe A 33 34 CNC 471506, 471507 1 A holotype 1323915749 CNCI, CNC Zimbabwe A 33 34 CNC 471515, 471516 10 10 A holotype 1323915765 [259,733,1519,1544] Australia Mt 33 34 1 Queensland holotype 1323915766 [744,1000,1519,1544] 1980-01-10 Australia Webb 33 34 2 2 Queensland holotype 1323915768 1980-03-10 ANIC J. C. Cardale Australia Rain forest 33 34 ANIC 32-070476, 32-070477 1 Queensland holotype 1323915761 1976-03-30 QDPC I. D. Galloway Australia Base Tinaroo Falls 100 Cableway Base Stn Bellenden Ker Range 33 34 1 Queensland holotype 1323915784 1981-09-11 QDPC Australia Bowenia State Forest 33 34 1 Cent holotype 1323915791 2005-12-17 Australia 30 Emmagen creek 145.45738 Bloomfield Track 33 34 1 1 Cape holotype 1323915772 2005-12-18 CNCI, CNC M. Buffington Australia Round Hill Head 33 34 CNC 471592 1 A holotype 1323915755 1975-05-05 QDPC I. Naumann & Mt. Finnigan Q. Australia Mt. Finnigan 33 34 1 A holotype 1323915777 1981-05-16 82-07-04 1981-05-16 ANIC I. D. Naumann Australia Gatton 33 34 ANIC 32-070480, 32 1 A holotype 1323915792 2002-12-17 CNCI, CNC J. Munro & Laceys Creek Q Rain forest Laceys Creek 34 35 CNC 471593 1 1 A 1323915769 1980-05-14 ANIC I. D. Naumann & J. C. Cardale 30 Mossman Gourge 34 35 ANIC 32-070489, 32- 070490 2 2 A 1323915771 QDPC Mt. Tamborine 34 35 1 1 A 1323915753 [672,974,404,429] 1981-04-27 Mt. Webb 34 35 4 4 A 1323915782 1974-04-27 QDPC Pebbly Beach, N & Cairns & J. F. Donaldson Mossman Rotary Park 34 35 1 1 A 1323915773 [259,599,548,573] 1980-10-17 Shiptons Flat 34 35 1 1 A 1323915793 1980-10-19 ANIC J. C. Cardale Townsville James Cook Uni Campus 34 35 ANIC 32-070499 1 A 1323915762 [586,1432,583,608] 1988-05-24 WINC A 34 35 1 A 1323915770 [151,1092,620,645] 1976-03-31 QDPC Tully Falls Road & I. D. Galloway Tully Falls 34 35 1 1 A 1323915783 1988-06-06 CALM Marun Western Australia 34 35 1 Western Australia 1323915781 1988-11-06 ANIC I. D. Naumann Mining Camp 34 35 ANIC 32-070500 3 3 Plateau 1323915779 03-07-05 1983-05-19 03-07-05 ANIC I. D. Naumann & J. C. Cardale Bangladesh BANGLADESH 34 35 ANIC 32-070501, 32-070502 1 1323915774 [151,1133,764,789] 1982-03-11 CNCI, CNC J. V. Harten Bangladesh Bangladesh 34 35 CNC 471531 1 1 1323915760 2002-06-22 SPC Bangladesh Compound 34 35 1 1 1323915775 2002-06-30 CNCI, CNC L. Leblanc French Polynesia Bora Bora 34 35 CNC 471532 1 1 1323915776 1984-04-09 CNCI, CNC D. M. Lasalle French Polynesia Bora Bora Tahitii 34 35 CNC 471533, 471534 3 3 1323915786 1982-07-21 CNCI, CNC H. Andersen Col. India INDIA 34 35 CNC 471536, 471537 1 1323915797 [252,752,908,933] CIBC Bangalore India Karnataka 34 35 1 1 Karnataka 1323915789 CNCI, CNC M. Hayat Indonesia INDONESIA 34 35 CNC 471540 1 1 1323915785 1978-11-21 CNCI, CNC Penulisan & J. T. Huber Indonesia Curug 34 35 CNC 471541 1 1 Bali 1323915809 1980-12-30 CNCI, CNC H. R. Gillespie & Neth. Guega, W. Indonesia 1200 Papua 34 35 CNC 471543, 471544 1 Papua 1323915822 1958-11-14 BPBM J. L. Gressitt Indonesia 220 Sieved 34 35 2 2 Sulawesi Utara 1323915794 1985-06-25 WINC Indonesia A 34 35 1 A 1323915778 1980-12-08 CNCI, CNC C. M. Yoshimoto Japan Mt. Tachibanayama 34 35 CNC 471545, 471546 1 Fukuoka 1323915802 [530,1083,1196,1221] Japan 1950 Takamine Table Land 34 35 1 1 Gunma 1323915817 1980-07-19 CNCI, CNC A. & Z. Smetana Japan Toyota Yakusa 34 35 CNC 471376 2 2 Aichi 1323915790 1990-07-08 CNCI, CNC K. Yamagishi Japan Tsuchiura 34 35 CNC 471272, 471273 1 Ibaraki 1323915796 [339,719,1303,1328] 1989-11-09 Japan Tsuchiura forest 34 35 1 1 Ibaraki 1323915780 1989-10-27 CNCI, CNC M. J. Sharkey Japan Tsuchiura 34 35 CNC 471280 1 Ibaraki 1323915805 1989-03-10 CNCI, CNC M. J. Sharkey Japan Shizizuka forest Tsuchiura 34 35 CNC 471281 2 2 Ibaraki 1323915801 [548,1430,1447,1472] 1996-05-09 CNCI, CNC L. Masner Japan Tsukuba 34 35 CNC 471282, 471283 6 6 Ibaraki 1323915816 1989-05-31 CNCI, CNC M. Sharkey Japan Matsushiro Tsukuba 34 35 CNC 471284, 471285 1 Ibaraki 1323915788 1989-04-08 CNCI, CNC M. J. Sharkey & Nishinasuno, N. G. R. I. Japan 500 Tochigi 34 35 CNC 471290 3 3 Tochigi 1323915795 1989-10-08 CNCI, CNC M. J. Sharkey Japan Mt. Tsukuba 34 35 CNC 471291, 471292 1 1 Ibaraki 1323915806 1989-06-09 CNCI, CNC M. J. Sharkey Japan Shishizuka-Oika Tsuchiura City 34 35 CNC 471295 1 1 Ibaraki 1323915808 1989-08-25 CNCI, CNC M. J. Sharkey Japan Tsuchiura 34 35 CNC 471296 1 1 Ibaraki 1323915803 1989-02-10 CNCI, CNC M. Sharkey Japan Expo Site Tsuchiura 34 35 CNC 471301 4 4 Ibaraki 1323915798 1989-08-08 CNCI, CNC M. J. Sharkey Japan Expo Site Tsukuba 34 35 CNC 471302, 471303 5 5 Ibaraki 1323915787 1989-03-11 CNCI, CNC M. Sharkey Japan Tsukuba 35 36 CNC 471351 1 Ibaraki 1323915799 1989-09-18 CNCI, CNC M. Sharkey Japan Mt Hiko Kyushu 35 36 CNC 471374 4 4 Fukuoka 1323915800 [295,1359,583,608] 1989-02-10 CNCI, CNC K. Takeno & M. Sharkey Japan Mt. Tachibana 35 36 CNC 471554 1 1 Fukuoka 1323915810 1992-08-17 CNCI, CNC K. Yamagishi Japan 1000 Nakakaruizawa 35 36 CNC 471563, 471564 4 4 Nagano 1323915818 1980-07-19 CNCI, CNC Nepal 2700 Kakani 35 36 CNC 471375 1 1 A 1323915813 CNCI, CNC Noumea & Ouen Toro & P. Cochereau New Caledonia NEW CALEDONIA 35 36 CNC 471571, 471572 1 1323915811 [278,939,908,933] 2006-01-08 New Caledonia 100 -4.266667 Morox 1306 144.96666 35 36 1 1 Madang 1323915832 2006-08-16 CNCI, CNC V. Iwam New Caledonia 5 -5.2833333 Nagada 1305 145.81667 35 36 CNC 471574 1 Madang 1323915838 2006-06-06 CNCI, CNC J. Hrcek New Caledonia 100 -5.25 Wanang 1305 145.26666 35 36 CNC 471575 1 Madang 1323915821 2006-08-23 CNCI, CNC P. Damen New Caledonia Morobe 35 36 CNC 471576, 471577 2 2 Morobe 1323915804 1983-07-31 CNCI, CNC S. & P. Miller New Caledonia Okapa 35 36 CNC 471578, 471579, CNC 471580 1 Eastern 1323915826 1964-10-27 ANIC R. Hornabrook New Caledonia South 35 36 ANIC 32-070505 1 South 1323915836 [394,669,1087,1112] Philippines Davao 35 36 1 Davao 1323915814 BMNH M. Gavarra Philippines Davao 35 36 1 1 Davao 1323915820 1987-07-15 Cuernos de Negros Philippines 700 9.283334 Cuernos de Negros 1300 123.25 35 36 1 1 Negros Oriental 1323915807 1987-07-29 CNCI, CNC D. C. Darling & E. Mayordo Singapore SINGAPORE 35 36 CNC 471582, 471583 1 1323915829 1978-11-30 CNCI, CNC J. T. Huber Iceland Honiara Guadalcanal 35 36 CNC 471528 1 1323915827 BPBM, CNCI, CNC N. L. H. Krauss & Munda Georgia 50 Georgia 35 36 CNC 471585 1 1323915834 2006-06-05 South Korea Chung-nam 36.400333 Wadong 11 127.433 Daejon-si 35 36 1 1 1323915815 2006-05-28 CNCI, CNC P. Tripotin South Korea Jirisan 700 35.34861 Macheon-myon 19 127.63917 Hamyang-gum 35 36 CNC 471377 2 2 1323915819 2002-09-14 CNCI, CNC P. Tripotin South Korea Kangwon Nam-myeon Chuncheon 35 36 CNC 471378, 471379 1 1 1323915831 2003-08-16 CNCI, CNC P. Tripotin South Korea On-dzong 450 Ok-ru Kus-gang 35 36 CNC 471381 1 1323915828 1975-05-08 HNHM J. Papp South Korea A 35 36 1 A 1323915830 1970-01-14 CNCI, CNC Kegalle & Mussard Besuchet Lobl South Korea Sabaragamuwa 35 36 CNC 471588 1 1 Sabaragamuwa 1323915823 1969-01-01 CNCI, CNC Nugegoda & B. P. Karumaratne Thailand 16.542433 PhetchabunKhao 35 36 CNC 471589 1 West 1323915812 2007-06-26 CNCI, CNC Somchai Chachumnan, S. Thailand 14.9429 Huai Mongpae 9 99.4205 35 36 CNC #2458 1 Suphanburi