Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species Ha, Tuyet Ngan Tran, Anh Duc European Journal of Taxonomy 2021 2021-06-24 756 1 82 Tran, 2021 Tran 2021 [549,941,1052,1079] Insecta Corixidae Micronecta GBIF Animalia Hemiptera 55 56 Arthropoda species nieseri sp. nov. Micronecta   urn:lsid:zoobank.org:act: 1AA8EB22-6D08-4894-81DA-E9CE0D648CFD  Figs 13G–H, 18    Diagnosis Body length 1.8–2.4. Pronotum longer than median head length. With dark and pale colour forms ( Fig. 13G–H). Males: fore femur with one pair of spines on proximal third ventrally, one longer spine on distal third dorsally, two spines distally, and one long seta dorsodistally; fore tibia with two long spines on distal third ventrally; palar claw distally moderately widened, apex rounded ( Fig. 18A). Median lobe of sternite VII with three or four long setae and an angular apex ( Fig. 18C). Free lobe short, distally strongly widened, posterior margin straight, lateral corner broadly notched, with a row of uniformly short, dark coloured setae restricted to lateral angle ( Fig. 18D). Left paramere: shaft relatively short and broad, with two ridges running to lateral margin, with the proximal ridge forming a triangular projection on lateral margin; mesial margin sinuate distally; apex tongue-shaped; basal lobe sub-triangular, directed towards paramere’s apex ( Fig. 18F). Right paramere: shaft with sinuate sides, abruptly constricted and bent at the middle part, apical part wavy and lobate ( Fig. 18E); pars stridens processus with ca 25 ridges.    Etymology This species is named after Dr Nico Nieser in honour of his great contributions to the taxonomy and our knowledge of Micronectidae.    Material examined   Holotype   VIETNAM• ♂; Lao Cai Prov., Sa Pa, Thanh Phu, Nam Cangstream;  26 Oct. 2013; A.D. Tran et al.leg.; TAD1359; ZVNU.   Paratypes   VIETNAM–  Lao Cai Prov.• 16 ♂♂, 4 ♀♀; same locality data as for holotype; ZVNU•  4 ♂♂, 2 ♀♀; same locality data as for holotype; NHMW•  7 ♂♂, 11 ♀♀; Sa Pa, Hoang Lien N.P., Cat Cat;  1 Jun. 2003; A.D. Tranleg.; TAD0336; ZRC•  3 ♂♂, 2 ♀♀; Sa Pa, Hoang Lien N.P., Sin Chai;  1 Jun. 2003; A.D. Tranleg.; TAD0337; ZRC•  1 ♂; Sa Pa Ban Den, Ta Van, Muong Hoastream;  24 Apr. 2011; N.H. Dinh et al.leg.; DNH11.13; ZVNU•  2 ♂♂, 1 ♀; Sa Pa, Nui Xe, Tram Ton area, tributary of Vang stream;  23 Apr. 2011; N.H. Dinh et al.leg.; DNH11.07; ZVNU•  2 ♂♂; Sa Pa, Ban Ho, Nam Pustream 1;  21 Oct. 2012; N.H. Dinh et al.leg.; DNH12.01; ZVNU•  11 ♂♂, 2 ♀♀; Sa Pa, Nam Cang, Nam Cangstream;  23 Oct. 2012; N.H. Dinh et al.leg.; DNH12.08; ZVNU•  1 ♀; Sa Pa, Nam Sai, Seo Nam Saistream 2;  24 Oct. 2012; N.H. Dinh et al.leg.; DNH12.10; ZVNU•  1 ♂, 3 ♀♀; Sa Pa, Ban Ho, Nam Pustream (tributary of Muong Hoa stream), site 1, at lower section;  26 Oct. 2013; A.D. Tran et al.leg.; TAD1361; ZVNU•  3 ♂♂; Sa Pa, Trung Chai, km 119 Nat. Road #4D;  28 Oct. 2013; Q.H. Ngo et al.leg.; TAD1370; ZVNU.    Description COLOUR ( Fig. 13G–H). With variations in colour of dorsum pattern of hemelytra, a dark form and a pale form. Most specimens with dark dorsum, generally darker in males ( Fig. 13G): frons and vertex brownish, eyes dark reddish brown. Pronotum dark brown (males) or light brownish yellow (females). Hemelytron generally light brownish, with ill-defined dark marks; in some specimens with minute reddish dots. Membrane translucent and generally brown. Venter of thorax and abdomen yellowish brown (males) or light yellow (females), legs pale yellow. Some specimens with pale, bright dorsum ( Fig. 13H): frons and vertex pale, eyes reddish brown. Pronotum sordid yellow but a little bit darker than vertex. Hemelytron without dark marks. Right membrane with similar colour and texture to corium. Left membrane generally hyaline. Venter pale yellow and legs very pale yellow or nearly whitish. Sample at locality TAD1359 with both colour forms.   Fig. 18.Morphological features of  Micronecta nieseri sp. nov., ♂, paratype (ZVNU). A. Fore leg. B. Prestrigilar flap. C. Median lobe of sternite VII. D. Free lobe. E. Right paramere. F. Left paramere.   Structural characteristics MACROPTEROUS MALES. Body length 2.13–2.44 ( holotype: 2.31), width 1.0–1.30 ( holotype: 1.25). Head width 0.82–1.01 ( holotype: 0.95). Pronotum well developed, about 2.4 times as wide as long (0.94: 0.40). Pronotum slightly wider than head. Synthlipsis 1.2 times as wide as posterior width of an eye (0.36: 0.29). Lengths of leg segments: fore leg: femur 0.38, tibia 0.20, pala 0.23; middle leg: femur 0.85, tibia 0.30, tarsus 0.36, claw 0.20; hind leg: femur 0.50, tibia 0.41, tarsus I 0.41, tarsus II 0.18, claw 0.07. Prestrigilar flap on segment V prolonged, with weakly acute apex ( Fig. 18B). Strigil present. Median lobe of sternite VII, free lobe, and parameres as in diagnosis. MACROPTEROUS FEMALES. Body length 1.84–2.3, width 0.8–1.0. Head width 0.88–0.9. Pronotum well developed, about 2.5 times as wide as long (0.94:0.37). Pronotum slightly wider than head. Synthlipsis 1.5 times as wide as posterior width of an eye (0.41: 0.27). Lengths of leg segments: fore leg: femur 0.32, tibiotarsus 0.3; middle leg: femur 0.81, tibia 0.38, tarsus 0.35, claw 0.25; hind leg: femur 0.59, tibia 0.32, tarsus I 0.4, tarsus II 0.2, claw 0.09.    Distribution  Vietnam: Lao Cai.    Remarks See Remarks under  M. pingae sp. nov. 3311572474 2013-10-26 ZVNU A. D. Tran Vietnam Sa Pa Nam Cang Thanh Phu 55 56 TAD1359 1 1 Lao Cai Prov. holotype 3311572478 [189,1219,299,326] ZVNU Vietnam Lao Cai Prov. 56 57 20 4 16 Lao Cai Prov. paratype 3311572343 NHMW Vietnam Lao Cai Prov. 56 57 6 2 4 Lao Cai paratype 3311572657 2003-06-01 ZRC A. D. Tran Vietnam Hoang Lien N. P. Sa Pa 56 57 TAD0336 18 11 7 Lao Cai paratype 3311572529 2003-06-01 ZRC A. D. Tran Vietnam Hoang Lien N. P. Sa Pa 56 57 TAD0337 5 2 3 Lao Cai paratype 3311572460 2011-04-24 ZVNU N. H. Dinh Vietnam Sa Pa Ban Den Muong Hoa Ta Van 56 57 DNH11.13 1 1 Lao Cai paratype 3311572422 2011-04-23 ZVNU N. H. Dinh Vietnam Sa Pa Tram Ton area Nui Xe 56 57 DNH11.07 3 1 2 Lao Cai paratype 3311572424 2012-10-21 ZVNU N. H. Dinh Vietnam Sa Pa Nam Pu Ban Ho 56 57 DNH12.01 2 2 Lao Cai paratype 3311572492 2012-10-23 ZVNU N. H. Dinh Vietnam Sa Pa Nam Cang Nam Cang 56 57 DNH12.08 13 2 11 Lao Cai paratype 3311572571 2012-10-24 ZVNU N. H. Dinh Vietnam Sa Pa Seo Nam Sai Nam Sai 56 57 DNH12.10 1 1 Lao Cai paratype 3311572356 2013-10-26 ZVNU A. D. Tran Vietnam Sa Pa Nam Pu Ban Ho 56 57 TAD1361 4 3 1 Lao Cai paratype 3311572519 2013-10-28 ZVNU Q. H. Ngo Vietnam Sa Pa Trung Chai Trung Chai 56 57 TAD1370 3 3 Lao Cai paratype