Takaoka, 2008b : 203 Classification, annotated list and keys for the black flies (Diptera: Simuliidae) of Peninsular Malaysia Takaoka, Hiroyuki Ya’Cob, Zubaidah Sofian-Azirun, Mohd Zootaxa 2018 2018-10-12 4498 1 1 65 Takaoka, 2008 Takaoka 2008 [193,781,1316,1341] Insecta Simuliidae Simulium GBIF Animalia Diptera 39 40 Arthropoda species charlesi Gomphostilbia     Simulium( Gomphostilbia) charlesi  Takaoka, 2008b: 203–210 (Female, male, pupa and larva). This species was described from females, males, pupae and larvae collected from Bario, Sarawak ( Takaoka, 2008b). It is characterized by the male style widened, and the pupal gill with eight filaments arranged as 3+3+2 from dorsal to ventral, of which the inner filament of the ventral pair is a little longer and thicker than the ourter filament.  This species was also collected from Peninsular Malaysia (Takaoka, 2012). The pupae and larvae of this species occur in small to medium-sized lowland streams.