Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae) WANG, YA-LIAN LI, BING-XUE CUI, MENG-DI ZHANG, YUAN WANG, LE ZHANG, CAI-HONG TSAUR, SHUN-CHERN CHEN, HONG-WEI HUANG, JIA Zootaxa 2023 2023-03-07 5250 1 1 109 BDMRS Zhang & Chen Zhang & Chen [151,730,1891,1918] Insecta Drosophilidae Stegana Animalia Diptera 95 96 Arthropoda species silvestrella sp. nov. Steganina  ( Figs. 13G, 13H, 61)  Diagnosis.This species is similar to S. ( S.) flavipedes sp. nov.in the shape of male terminalia (Figs. 38, 61), but can be distinguished from the latter by having the pregonite blurry or absent ( Fig. 61C, D); the aedeagus with numerous minute, serrated processes distally ( Fig. 61C, D).  Description.Male and female: First flagellomere yellow. Clypeus yellow. Mesonotum brown medially, brownish yellow laterally ( Fig. 13G). Katepisternum yellowish ( Fig. 13H). Scutellum dark brown in male, brownish yellow in female, black near distal margin ( Fig. 13G). Halter mostly yellowish ( Fig. 13H). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites brown medially, yellow laterally ( Fig. 13G, H); sternites yellow, about 1.8 times as wide as long. Male terminalia ( Fig. 61): Epandrium with ca. 9 setae on dorsal to posterior portions per side ( Fig. 61A). Surstylus with some setulae near distal margin, lacking serrated processes and pubescence ( Fig. 61A, B). Cercus with much pubescence ( Fig. 61A). Hypandrium roundly expanded anteromedially ( Fig. 61C, D). Aedeagus tubular, distally expanded and rounded, lacking pubescence ( Fig. 61C, D). Aedeagal sheath lacking serrated process ( Fig. 61C, D). Subepandrial sclerite slightly half-rounded ( Fig. 61G).   FIGURE 61.  Stegana( Steganina) silvestrellaZhang & Chen,  sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view.  Measurements and indices.BL = 2.63 mmin holotype( 2.80–3.40 in 2♀ paratypes), ThL = 1.33 mm( 1.33 in ♀), WL = 2.13 mm(2.00– 2.47 in ♀), WW = 1.00 mm (1.00– 1.13 in ♀), arb = 4/3 (4–5/3–4), avd = 1.00 (0.80–1.14), adf = 1.25 (1.00–1.40), flw = 2.42 (2.01–2.50), FW/HW = 0.45 (0.39–0.53), ch/o = 0.09 (0.08–0.11), prorb = 1.38 (0.83–1.33), rcorb = 0.75 (0.50–0.78), vb = 0.67 (0.60–0.80), dcl = 0.46 (0.46–0.48), presctl = 0.72 (0.70–0.82), sctl = 2.34 (2.11–2.50), sterno = 0.80 (0.60–1.11), orbito = 2.67 (2.12–2.67), dcp = 0.28 (0.25–0.28), sctlp = 1.86 (1.75–2.50), C = 1.67 (1.56–2.00), 4c = 1.29 (1.13–1.50), 4v = 1.91 (1.43–2.01), 5x = 2.43 (2.01–2.60), ac = 10.02 (9.00–12.01), M = 0.76 (0.73–0.84), C3F = 0.59 (0.64–0.87).  Type materials.  Holotype♂ ( SCAU, No. 125657, DNA #2752), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt.  700m,  25.iv.2007, ex tree trunk, H.W. Chen.  Paratypes: 2♀( SCAU, Nos. 125658, 59, DNA#2562), Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt.  1320m,  10.v.2016, ex tree trunks, J. Huang, Y.L. Wang.   Distribution. China( Yunnan).   Etymology.From the Latinword " silvestris" (= arboreal), referring to the arboreal  Steganinaflies. 4097216469 2007-04-25 SCAU H. W. Chen China 700 97 98 23.168888 Wangtianshu 21 99.23111 96 97 1 Yunnan holotype 4097216485 2016-05-10 SCAU J. Huang China 1320 23.168888 Mengdong 21 99.23111 97 98 2 2 Yunnan paratype