Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae) WANG, YA-LIAN LI, BING-XUE CUI, MENG-DI ZHANG, YUAN WANG, LE ZHANG, CAI-HONG TSAUR, SHUN-CHERN CHEN, HONG-WEI HUANG, JIA Zootaxa 2023 2023-03-07 5250 1 1 109 BDMST Li & Chen Li & Chen [151,640,294,321] Insecta Drosophilidae Stegana Animalia Diptera 68 69 Arthropoda species flavipes sp. nov. Steganina  ( Figs. 8E, 8F, 38)   Diagnosis.This species is similar to S. ( S.) silvestrella  sp. nov.in the shape of male terminalia ( Figs. 38, 61), but can be distinguished from the latter by having the small pregonite with 2 sensilla ( Fig. 38C, D); the aedeagus lacking serrated process ( Fig. 38C, D).   Description.Male and female: First flagellomere brown. Clypeus brownish yellow medially, yellow laterally. Mesonotum brown medially, yellow laterally ( Fig. 8E). Katepisternum yellowish ( Fig. 8F). Scutellum dark brown to black ( Fig. 8E). Halter mostly greyish ( Fig. 8F). Fore femur with a row of 4 thin, long setae on distal part of posteroventral surface. Abdominal tergites dark brown medially, yellow laterally ( Fig. 8E, F); sternites yellow, about 1.6 times as wide as long. Male terminalia ( Fig. 38): Epandrium with ca. 8 setae on dorsal to posterior portions per side ( Fig. 38A). Surstylus slightly oval, with some setulae along distal margin and inner surface, lacking pubescence ( Fig. 38A, B). Cercus with sparse pubescence ( Fig. 38A). Hypandrium expanded anteromedially ( Fig. 38C, D). Aedeagus tubular, slightly expanded distally, lacking serrated processes and pubescence ( Fig. 38C, D). Aedeagal sheath small, lacking serrated process ( Fig. 38E, F). Subepandrial sclerite broadened, slightly squared ( Fig. 38G).   FIGURE 38.  Stegana( Steganina) flavipesLi & Chen,  sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F. Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view.  Measurements and indices.BL = 2.23 mmin holotype( 2.24–2.92 in5♂, 2.19–2.61 in 5♀ paratypes), ThL = 1.23 mm( 1.08–1.43 in♂, 1.16–1.35 in ♀), WL = 2.21 mm( 2.45–2.79 in♂, 1.91–2.41 in ♀), WW = 1.21 mm( 0.93–1.31 in♂, 0.87–1.01 in ♀), arb = 6/4 (6–7/4–5), avd = 0.82 (0.61–0.90), adf = 0.95 (0.94–1.50), flw = 1.76 (1.33–2.12), FW/HW = 0.39 (0.35–0.44), ch/o = 0.32 (0.26–0.41), prorb = 0.72 (0.58–1.49), rcorb = 0.96 (0.54– 1.04), vb = 0.82 (0.49–1.14), dcl = 0.59 (0.29–0.62), presctl = 0.71 (0.45–0.82), sctl = 1.65 (1.52–3.02), sterno = 1.45 (0.61–1.10), orbito = 1.73 (1.65–2.24), dcp = 0.24 (0.17–0.26), sctlp = 1.71 (1.39–2.00), C = 2.13 (1.63–2.01), 4c = 1.11 (1.09–1.31), 4v = 1.82 (1.77–2.18), 5x = 1.72 (1.21–1.86), ac = 8.79 (9.33–12.11), M = 0.70 (0.50–0.79), C3F = 0.71 (0.53–0.69).   Type materials.  Holotype♂ ( SCAU, No. 125671, DNA #1303), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt.  660m,  30.ix.2011, ex tree trunk, H.W. Chen.  Paratypes: 1♂, 2♀( SCAU, Nos.125655, 72, 73, DNA #2686, 2746), 1♂ on  25.iv.2007,  2♀on  30.ix.2011, other data same as holotype; 1♂, 2♀( SCAU, Nos.125674–76, DNA #2171), Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt.  1320m,  10.v.2016, ex tree trunks, J. Huang; 4♂, 5♀( SCAU, Nos.125677–85, DNA #2684), Longtan Park, Ximeng, Yunnan, 22°38'55''N, 99°36'02''E, alt.  1150m,  17.iv.2018, ex tree trunks, L. Gong, Y. Lin.   Distribution. China( Yunnan).   Etymology.A combination of the Latin words " flavus" (= yellow) and "- pes" (= leg), referring to the yellow legs. 4097216355 2011-09-30 SCAU H. W. Chen China 660 21.6025 Mengla 21 101.585 Wangtianshu 69 70 1 Yunnan holotype 4097216362 2007-04-25 SCAU China Nos. 69 70 2 2 Yunnan paratype 4097216400 [668,1191,367,393] 2011-09-30 SCAU H. W. Chen China 660 21.6025 Mengla 21 101.585 Wangtianshu 69 70 2 2 Yunnan paratype 4097216369 2011-09-30 SCAU H. W. Chen China Nos. 660 21.6025 Mengla 21 101.585 Wangtianshu 69 70 2 2 Yunnan paratype 4097216479 2016-05-10 SCAU J. Huang China Nos. 1320 23.168888 Cangyuan 21 99.23111 Mengdong 69 70 2 2 Yunnan paratype