Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae) WANG, YA-LIAN LI, BING-XUE CUI, MENG-DI ZHANG, YUAN WANG, LE ZHANG, CAI-HONG TSAUR, SHUN-CHERN CHEN, HONG-WEI HUANG, JIA Zootaxa 2023 2023-03-07 5250 1 1 109 Li & Chen Li & Chen [151,642,870,897] Insecta Drosophilidae Stegana Animalia Diptera 60 61 Arthropoda species cornuta sp. nov. Steganina  ( Figs. 7A, 7B, 32)   Diagnosis.This species closely resembles S. ( S.) angulistrata Wu & Chen, 2010in the shape of male terminalia (as fig. 1–6 in Wu & Chen 2010, Fig. 32), but can be distinguished from the latter by having the aedeagus medially lacking serrated process but mediolaterally with a pair of horned processes ( Fig. 32C, D); the aedeagal sheath distally slender and strongly dorsad curved ( Fig. 32E, F).   Description.Male and female: First flagellomere brown. Clypeus yellow. Mesonotum yellow ( Fig. 7A). Katepisternum yellowish ( Fig. 7B). Scutellum yellow ( Fig. 7A). Halter mostly grey ( Fig. 7B). Fore femur with a row of 3 or 4 long setae on distal part of posteroventral surface. Abdominal tergites brown to dark brown ( Fig. 7A, B); sternites brown, slightly wider than long. Male terminalia ( Fig. 32): Epandrium with ca. 9 setae on dorsal to posterior portions per side ( Fig. 32A). Surstylus with 1 prensiseta, several setulae, and serrated processes near distal margin ( Fig. 32A, B). Cercus lacking pubescence ( Fig. 32A). Hypandrium expanded anteromedially ( Fig. 32C, D). Pregonite small, with 2 sensilla ( Fig. 32C, D). Aedeagus rounded distally, with numerous minute, serrated processes laterally and much pubescence distoventrally and sublaterally ( Fig. 32C, D). Aedeagal sheath with some minute, serrated processes distally ( Fig. 32E, F). Subepandrial sclerite slightly vaulted, split medially ( Fig. 32G).  Measurements and indices.BL = 3.00 mm in holotype( 2.34–3.04 in4♂, 2.88–3.44 in 5♀ paratypes), ThL = 1.48 mm( 1.21–1.46 in♂, 1.29–1.80 in ♀), WL = 2.32 mm( 2.31–2.59 in♂, 2.43–2.72 in ♀), WW = 1.20 mm( 0.97–1.09 in♂, 0.91–1.24 in ♀), arb = 9/5 (6–9/4–6), avd = 0.83 (0.76–0.93), adf = 1.50 (0.91–1.29), flw = 2.50 (1.74–2.16), FW/HW = 0.48 (0.27–0.46), ch/o = 0.17 (0.26–0.37), prorb = 1.50 (0.53–1.41), rcorb = 0.70 (0.59– 1.16), vb = 0.56 (0.46–1.22), dcl = 0.33 (0.43–0.69), presctl = 0.57 (0.39–0.63), sctl = 2.23 (1.47–2.27), sterno = 0.64 (0.53–0.96), orbito = 1.40 (1.67–1.98), dcp = 0.28 (0.16–0.39), sctlp = 1.89 (1.55–2.08), C = 1.74 (1.53–2.13), 4c = 1.55 (1.06–1.38), 4v = 2.40 (1.36–2.10), 5x = 1.50 (1.06–2.00), ac = 15.50 (8.36–17.55), M = 0.75 (0.58–0.72), C3F = 0.81 (0.56–0.71).   Type materials.  Holotype♂ ( SCAU, No. 125586, DNA #1414, 2665), CHINA: Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt.  1420m,  13.v.2012, ex tree trunk, H.W. Chen.  Paratypes: 2♂, 5♀( SCAU, Nos. 125587–93, DNA #1414, 1939, 2665, 2702), 1♂, 1♀on  28–30.iii.2013, J.J. Gao, other data same as holotype; 1♂ ( SCAU, No. 125594, DNA #2173), Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt.  1320m,  10.v.2016, ex tree trunk, J. Huang.   Distribution. China( Yunnan). 4097216432 2012-05-13 SCAU H. W. Chen China 1420 22.738611 Pu'er 21 101.11694 Yixiang 60 61 1 Yunnan holotype 4097216460 SCAU China 60 61 5 5 Yunnan paratype 4097216426 2013-03-28 2016-05-10 2013-03-28 SCAU J. J. Gao & J. Huang China 1320 23.168888 Cangyuan 21 99.23111 Mengdong 60 61 5 5 Yunnan paratype