Illustrated checklist of the carabid beetles (Coleoptera, Carabidae) of the Otonga Nature Reserve in biogeographic comments NW Ecuador, with ecological and Moret, Pierre Allegro, Gianni Faunitaxys 2025 New York, N. Y. 13 17 1 21 W62H Grouvelle, 1903 Grouvelle 1903 [992,1353,1289,1312] Insecta Carabidae Clinidium Animalia Coleoptera 5 3 Arthropoda species insigne  ( Fig. 3) Rotten logs, under barks and in decomposed wood. March, July, November (CPMo, QCAZ).  Other provinces:  Pichincha, Bolívar( Bell& Bell, 2000).  Other countries: Colombia, Venezuela(Bell & Bell, 1985; Bell & Bell, 2009). Brachypterous. [1017,1433,1417,1440] Bell Pichincha, Bolivar 5 3 1 Pichincha, Bolivar