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        <cito:cites>Psammoecus obesus Grouvelle, 1919: 11</cito:cites>
        <cito:cites>Psammoecus obesus</cito:cites>
        <cito:cites>Hetschko 1930: 83</cito:cites>
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        <dc:title>Taxonomic Studies on Australian Psammoecus Latreille (Coleoptera, Silvanidae, Brontinae)</dc:title>
        <dc:creator>Karner, Michael</dc:creator>
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        <bibo:journal>European Journal of Taxonomy</bibo:journal>
        <dc:date>2020</dc:date>
        <bibo:pubDate>2020-11-12</bibo:pubDate>
        <bibo:volume>723</bibo:volume>
        <bibo:pageStart>135</bibo:pageStart>
        <bibo:pageEnd>158</bibo:pageEnd>
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        <dwc:authority>Grouvelle, 1919</dwc:authority>
        <dwc:authorityName>Grouvelle</dwc:authorityName>
        <dwc:authorityYear>1919</dwc:authorityYear>
        <dwc:box>[571,1016,581,608]</dwc:box>
        <dwc:class>Insecta</dwc:class>
        <dwc:family>Silvanidae</dwc:family>
        <dwc:genus>Psammoecus</dwc:genus>
        <dwc:kingdom>Animalia</dwc:kingdom>
        <dwc:order>Coleoptera</dwc:order>
        <dwc:pageId>7</dwc:pageId>
        <dwc:pageNumber>142</dwc:pageNumber>
        <dwc:phylum>Arthropoda</dwc:phylum>
        <dwc:rank>species</dwc:rank>
        <dwc:species>obesus</dwc:species>
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        <spm:hasContent>  Fig. 3</spm:hasContent>
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        <spm:hasContent>     Psammoecus obesus Grouvelle, 1919: 11.    Psammoecus obesus–  Hetschko 1930: 83.</spm:hasContent>
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        <spm:hasContent>   Diagnosis The following combination of character states distinguishes this species: body elongate-oval ( Fig. 3A), length 3.08–3.29 mm; eyes large ( Fig. 3B), irregularly rounded with stronger curvature posteriorly; temples distinct, short, strongly curved; frontal grooves distinct, diverging anteriorly, parallel near end; reaching anterior ²⁄5of eyes; antennae stout ( Fig. 3D), antennomere 8 slightly darkened, antennomeres 9–10 darkened, antennomere 11 bright yellowish-brown; punctation on vertex moderate, punctures distinctly elongate, as large as eye facet diameter; microsculpture on vertex absent, pubescence moderate; pronotum widest just in front of middle ( Fig. 3C), lateral pronotal margins with five teeth; tooth I slightly and tooth V distinctly bigger than II–IV, distance between I and II wider than distances between other teeth, anterior and posterior denticles present, small; pronotal punctation dense, punctures as large as on vertex, distinctly widened, microsculpture absent, pubescence moderate, setae directed medially; elytra widest at middle ( Fig. 3A), with transverse brown maculae behind middle, interstices distinctly wider than striae, pubescence moderate, microsculpture absent; male genitalia ( Fig. 3E–H) strongly sclerotised, median lobe in dorsal view wide, narrowed abruptly near ostium, apical end well rounded, in lateral view also narrowed abruptly, with sharp apical end; parameres in ventral view very wide, inner margins, apex and apical third of lateral margins with distinct setae; in lateral view with very wide basis, narrowed abruptly towards apex with dorsal margin almost forming angle.</spm:hasContent>
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        <spm:hasContent>   Material examined   Holotype  INDONESIA• ♀; Maluku, Seram; Pascoeleg.; “HOLO - | Type” [round label with red border]; “Ceram” [blue, elliptic label]; “Pascoe | Coll. | 93–60”; “  Psammoecus|  obesus| ty. Grouv” [Grouvelle’s hand]; “HOLOTYPUS |  Psammoecus|  obesusGrouv. 1919| det. R.G. Booth2013”; NHMUK.   Other material  AUSTRALIA• 1 spec.; Queensland, Cairns; 16°55′ S, 145°46′ E; 1942; Perkinsleg.; NHMUK•  1 spec.; Queensland, Cassowary Coast Region,  10 kmNE of Tully;  Nov. 1994; C. Reidleg.; rainforest; ANIC•  1 spec.; Queensland, Cassowary Coast Region, Cardstone;  1–2 Dec. 1966; J.G. Brooksleg.; ANIC•  1 spec.; Queensland, Shire of Cook, McIlwraith Range;  500 ma.s.l.; 13°45′ S, 143°22′ E;  27 Jun.–12 Jul. 1989; T.A. Weirleg.; at light; ANIC•  1 spm; Queensland, Shire of Douglas, Daintree;  29 Oct. 1966; E. Brittonleg.; dead leaves, edge of rainforest; ANIC•  1 spm; same data as for preceding; MKF.   INDONESIA• 2 specs; West Papua, Waigeo, Camp Nok;  2500 fta.s.l.; 0°12′ S, 130°50′ E;  Apr. 1938; Cheesmanleg.; NHMUK.</spm:hasContent>
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        <spm:hasContent>   Distribution The species has been found in Australia( Queensland) and Indonesia.</spm:hasContent>
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        <spm:hasContent>   Remarks   Psammoecus obesusis closely related to  P. hirsutusGrouvelle, 1883,  P. decoratus Grouvelle, 1919, and  P. signatus Grouvelle, 1919. It differs from all three by the shape of the male genitalia.</spm:hasContent>
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        <dwc:collectorName>Pascoe</dwc:collectorName>
        <dwc:country>Indonesia</dwc:country>
        <dwc:determinerName>R. G. Booth</dwc:determinerName>
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        <dwc:collectorName>Perkins</dwc:collectorName>
        <dwc:country>Australia</dwc:country>
        <dwc:latitude>-16.916666</dwc:latitude>
        <dwc:location>Cairns</dwc:location>
        <dwc:longLatPrecision>1280</dwc:longLatPrecision>
        <dwc:longitude>145.76666</dwc:longitude>
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        <dwc:collectedFrom>rainforest</dwc:collectedFrom>
        <dwc:collectingDate>1994-11</dwc:collectingDate>
        <dwc:collectionCode>ANIC</dwc:collectionCode>
        <dwc:collectorName>C. Reid</dwc:collectorName>
        <dwc:country>Australia</dwc:country>
        <dwc:location>Cassowary Coast Region</dwc:location>
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        <dwc:box>[209,1381,1776,1803]</dwc:box>
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        <dwc:collectingDateMax>1966-12-02</dwc:collectingDateMax>
        <dwc:collectingDateMin>1966-12-01</dwc:collectingDateMin>
        <dwc:collectionCode>ANIC</dwc:collectionCode>
        <dwc:collectorName>J. G. Brooks</dwc:collectorName>
        <dwc:country>Australia</dwc:country>
        <dwc:location>Cardstone</dwc:location>
        <dwc:municipality>Cassowary Coast Region</dwc:municipality>
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        <dwc:collectorName>T. A. Weir</dwc:collectorName>
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        <dwc:elevation>500</dwc:elevation>
        <dwc:latitude>-13.75</dwc:latitude>
        <dwc:location>McIlwraith Range</dwc:location>
        <dwc:longLatPrecision>1289</dwc:longLatPrecision>
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        <dwc:municipality>Shire of Cook</dwc:municipality>
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        <dwc:collectingDate>1966-10-29</dwc:collectingDate>
        <dwc:collectionCode>ANIC</dwc:collectionCode>
        <dwc:collectorName>E. Britton</dwc:collectorName>
        <dwc:country>Australia</dwc:country>
        <dwc:location>Daintree</dwc:location>
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        <dwc:collectionCode>MKF</dwc:collectionCode>
        <dwc:collectorName>E. Britton</dwc:collectorName>
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        <dwc:location>Daintree</dwc:location>
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        <dwc:collectionCode>NHMUK</dwc:collectionCode>
        <dwc:collectorName>Cheesman</dwc:collectorName>
        <dwc:country>Indonesia</dwc:country>
        <dwc:elevation>762</dwc:elevation>
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        <dwc:location>Camp Nok</dwc:location>
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