Harpalus spadiceus Harpalus comis Harpalus carolinae Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico Bousquet, Yves ZooKeys 2012 2012-11-28 245 1 1722 6LDT5 03671C90-F9D4-16C4-A2CB-4E5FC33786AA Dejean, 1829 Dejean 1829 Insecta Carabidae Harpalus GBIF Animalia Harpalus spadiceus Coleoptera 698 699 Arthropoda species spadiceus   Harpalus spadiceusDejean, 1829: 336. Type locality: "Ameriqueseptentrionale" (original citation), restricted to "White Sulphur Springs [Greenbrier County], W[est] Virg[ina]" by Lindroth (1968: 780). Holotype [by monotypy] (♀) in MHNP (Lindroth 1955b: 27).  Harpalus comisHaldeman, 1843b: 301 [ nomen dubium]. Type locality: southeastern Pennsylvania (Haldeman 1843a: 297). Syntype(s) presumably lost. Synonymy established with doubt by LeConte (1863b: 13).  Harpalus carolinaeSchaeffer, 1910: 402. Type locality: "Black Mountains, North Carolina" (original citation). Lectotype (♂), designated by Erwin and House (1978: 238), in USNM [# 42509]. Synonymy established by Noonan (1991: 86).  Distribution. This eastern species ranges from southern Quebec, southern Ontario, and the New England states south through the Appalachian mountains to northern Georgia [see Noonan 1991: Figs 273-275]. The record from southwestern Manitoba (Stjernberg 2011: 70) is probably in error.  Records.  CAN: ON, QC USA: CT, DC, DE, GA, KY, MA, NC, NH, NJ, NY, OH, PA, RI, SC, TN, VA, VT, WV  Note. Noonan (1991: 87-88) recognized two morphs for this species, the spadiceusand carolinaemorphs, that Lindroth (1968: 780-781) treated as distinct species. According to Noonan (1991: 88), the two morphs intergrade over an area extending southwards from Virginia to Georgia.