Paragymnopleurus, Shipp, 1897

Pokorný, Svatopluk & Zídek, Jiri, 2014, Review of the Gymnopleurini (Coleoptera: Scarabaeidae: Scarabaeinae). II. The genus Paragymnopleurus Shipp, Insecta Mundi 2014 (395), pp. 1-10 : 2-3

Checklist of Paragymnopleurus

Paragymnopleurus Shipp, 1897: 166 , type sp. Scarabaeus sinuatus Olivier , by orig. des. Syn. Progymnopleurus Garreta, 1914: 52 , type sp. Scarabaeus sinuatus Olivier (see Janssens 1940: 5).

aethiops ( Sharp, 1875: 34); HT – MNHN; = melanarius . ambiguus Janssens, 1943: 1 ; HT+3PT – ISNB; Taiwan. Fig. 1 brahminus ( Waterhouse, 1890: 411) ; HT – BMNH; China (Fujian, Shanghai, Sichuan, Zhejiang [npr –

JZPC, OXUM]), n. Vietnam (Vind Phu Prov.: Tam Dao; ncr – SPPC). Fig. 3 calcar ( Sharp, 1875: 36); HT – MNHN; = planus . celebicus ( Sharp, 1875: 37); HT – MNHN; = planus . dubius ( Sharp, 1875: 36); HT – MNHN; = planus . leei ( Donovan, 1798: pl. 1.4), as Scarabaeus ; type lost; = sinuatus (syn. by Janssens 1940: 20). martinezi Balthasar, 1955: 393 ; HT – NMPC; China (Sichuan); [see Comments]. Fig. 4 , 21 maurus maurus ( Sharp, 1875: 34) ; HT – MNHN; Malaysia, Borneo, Sumatra. Fig. 5 maurus malayanus Ochi and Kon, 1997: 236; HT+14PT – NSMT; Malaysia, w. Sumatra; = m. maurus ,

syn. n. [see Comments]. Fig. 6 maurus pauliani Janssens, 1940: 17 +19, as sp.; HT+1PT – ISNB; Borneo ( Sarawak; Brunei, ncr –

SPPC); = m. maurus , syn. n. [see Comments]. Fig. 7 melanarius ( Harold, 1867: 76) ; HT – MNHN; China ( Hong Kong, Shanghai, Sichuan), Taiwan, India,

Sri Lanka, Laos, Vietnam, Borneo, Java, Sumatra. Fig. 8 , 23 planus ( Sharp, 1875: 35) ; HT – MNHN; Malaysia, Sulawesi. Fig. 9 rudis ( Sharp, 1875: 37) ; HT – MNHN; Flores, Java, Lombok, Sumbawa, Thailand. Fig. 10 singularis ( Waterhouse, 1890: 410) ; “ Corea ”; LTm+2PLT – BMNH; = maurus [see Comments]. Fig. 11 sinuatus sinuatus ( Olivier, 1789: 160) ; HT – MNHN; s. China, Taiwan, Korean peninsula, Nepal, India,

se. Asia, Indonesia. Fig. 12 sinuatus abax ( Sharp, 1875: 39) , as sp.; HT – MNHN; Cambodia. sinuatus assamensis ( Waterhouse, 1890: 411), as sp.; HT – BMNH; n. India, Nepal, China, Myanmar.

Fig. 13 sinuatus productus ( Sharp, 1875: 38) , as sp.; HT – MNHN; Laos (HT), Cambodia, Vietnam. sinuatus szechouanicus Balthasar, 1934: 149; HT+2PT – NMPC; China (Sichuan, Xizang), Thailand

(ncr – JZPC). Fig. 14 sparsus sparsus ( Sharp, 1875: 38) ; HT – MNHN; Malaysia, s. Kalimantan. Fig. 15 sparsus arnoldi Krikken and Huijbregts, 1987: 3; HT – RMNH; Sumatra. Fig. 16 sparsus javanus Krikken and Huijbregts, 1987: 3; HT – RMNH; Java. Fig. 17 sparsus palawanicus Ochi and Kon, 1997: 236; HT+5PT – NSMT; Philippines (Palawan); = s. sparsus

[see Comments]. spinotus ( Boucomont, 1914: 248) , as var. of G. maurus ; HT – MNHN; Borneo ( Kina Balu ); = striatus

[var. of striatus in Janssens 1940: 18]. Fig. 18 stipes stipes ( Sharp, 1875: 35) ; HT – MNHN; Philippines (Mindoro). stipes japonicus Balthasar, 1955: 395, as ssp. (as var. on label); HT – NMPC; Japan (Honshu); = ambiguus ,

syn. n. [see Comments]. Fig. 2 striatus ( Sharp, 1875: 33) ; HT – MNHN; Singapore, Borneo, Sumatra. Fig. 19 sumatrensis Ochi and Kon, 1997: 235 ; HT+7PT – NSMT; w. Sumatra. Fig. 20

Figures

Figures 1-9. 1) Paragymnopleurus ambiguus Janssens, female, 23 mm. 2) P. stipes japonicus Balthasar [= P. ambiguus], male holotype, 21 mm. 3) P. brahminus (Waterhouse), female, 21 mm. 4) P. martinezi Balthasar, male holotype, 14 mm. 5) P. maurus maurus (Sharp), male, 16.5 mm. 6) P. maurus malayanus Ochi and Kon, female, 15.5 mm. 7) Paragymnopleurus maurus pauliani Janssens, female, 15.5 mm. 8) P. melanarius (Harold), male, 19.5 mm. 9) P. planus (Sharp), male, 17 mm.

Figures 19-24. 19) Paragymnopleurus striatus (Sharp), male, 20 mm. 20) P. sumatrensis Ochi and Kon, female, 16 mm. 21) Clypeus of P.martinezi. 22) Mesotibial spurs of P. sumatrensis. 23) Rounded lateral margin of pronotum (P. melanarius). 24) Angular lateral margin of pronotum (P. brahminus).

Figures 10-18. 10) P. rudis (Sharp), male, 18 mm. 11) P. singularis (Waterhouse) [= P. maurus], male lectotype, 19 mm. 12) P. sinuatus sinuatus (Olivier), male, 17 mm. 13) Paragymnopleurus sinuatus assamensis (Waterhouse), male, 15 mm. 14) P. sinuatus szechouanicus Balthasar, female, 20 mm. 15) P. sparsus sparsus (Sharp), female, 15 mm. 16) P. sparsus arnoldi Krikken and Huijbregts, male, 13 mm. 17) P. sparsus javanus Krikken and Huijbregts, female, 15 mm. 18) P. spinotus (Boucomont), female, 19 mm.