Glyphotmethis efe Ünal, 2007

Ünal, Mustafa, 2007, Revision of the genus Glyphotmethis Bey-Bienko, 1951 (Orthoptera: Pamphagidae), Zootaxa 1581, pp. 1-36 : 29-31

Glyphotmethis efe Ünal , sp.n.

( Figs. 13, 15; Table 1)

Eremobia Escherichi : Kuthy 1907: 432. (?)

Tmethis escherichi: Uvarov 1930: 372 .

Glyphotmethis escherichi escherichi: Cejchan 1965: 470 , 472.

Asiotmethis escherichi elatior: Ramme 1951: 274 .

Type locality: Turkey: Manisa, Selendi. Holotype: male ( AIBUEM).

Previous records. Kuthy 1907 (?): as Eremobia Escherichi : Bos-Dagh, Kara-Tschehir, July and August 1906, leg. A. Lendl (not examined, see Remarks).

Material examined: 35 specimens. TURKEY: Manisa, Selendi, Simav yolu, 1150 m, 38°51’996” K, 28°55’727” D, 6.6.2002 , 3 ♂, 2 ♀ (including holotype), leg. M. Ünal & S. Mutun ( AIBUEM); Manisa, Selendi, Kürkçü , 1940 , 1 ♂, 1941, 1 ♀ ( NHM); Kürkçü , 17.6.1940 , 3 ♀ ( AUZM); Kürkçü , 1 ♂ ( ZMMAE); Kürkçü, August , 1941 , 2 ♂, 3 ♀; August , 1946 , 1 ♂; Manisa, Salihli , 20.7.1941 , 2 ♀ ( AUZM); Uschak [Uşak], 2.6.1916 , 2 ♀, leg. S.G. Bauer ( NHM); Uschak , 2.6.1916 , 1 ♀, leg. S.G. Bauer (paratype of G. escherichi elatior ) ( ZMB); Uşak, Banaz , 910 m, 19.6.2001 , 1 ♂, 1 ♀, leg. M. Ünal & S. Mutun ( AIBUEM); Kütahya, 5.7.1930 , 2 ♂; Kütahya, Demirci , 1929 , 1 ♂, leg. Sureya ( NHM); Kütahya, Gediz, Çavdarhisar yolu 8 km, 890 m, 19.6.2001 , 4 ♂, 4 ♀, leg. M. Ünal & S. Mutun ( AIBUEM) .

Description and Diagnosis: (Holotype: male): Body small and slender; pronotum ( Figs. 13a, c) with distinct tubercles; prozona high with distinct lobes, incision between the lobes deep; lateral projections of mesozona triangular, large and sharp; metazona narrow and short, slightly raised in some specimens, lateral margins straight, hind margin acute angular; male tegmina ( Figs. 13a, c, e) short, cover the first 3 abdominal tergites from above, its widest point in middle; axillary 1 very indistinct, touching to axillary 2 many times and fused to it towards apex; distal excision of dorsal carina of hind femur less distinct, its ventral carina slightly wavy; inner side of hind femur very light pale red (pinkish-red) with a small distinct dark macula, its ventral and apical parts light pale red; inner side of hind tibia light pale red; male penis valves ( Figs. 13g, h) slender apically, their apices very long and thin, strongly curved upwards in lateral view, strongly narrowed in posterior view; lophi ( Fig. 13i) with less than 10 sparse spines in typical form, posterior margin of epiphallus ( Fig. 13i) with a slightly rounded projection. Female: Pronotum ( Figs. 13b, d) with distinct tubercles; prozona high and long, its lobes distinct; lateral projections of mesozona distinct; metazona flat, narrow and short, only 1.1 times longer than prozona, 1.5 times wider than long; its hind margin angular, slightly rounded in some specimens; tegmina ( Figs. 13b, d, f) pointed at apex, reaching to half of second abdominal tergite; distal excision of dorsal carina of hind femur less distinct, its ventral carina slightly undulating; inner side of hind femur red with a small dark macula in basal part dorsally; inner side of hind tibia light pale red.

This new species is unique in the structure of male penis valves, very thin and strongly prolonged upwards. It can be compared with the following taxa: This species has similar metazona of pronotum and hind femur (with less distinct apical excision of dorsal carina and its less wavy ventral carina) with G. heldreichi . It is different from it by the shape of penis valves, shorter prozona, larger lateral projections of mesozona, mostly flat metazona, indistinct axillary 1 vein; color of hind legs (yellow in G. heldreichi ); slender body and distribution. It is near to G. esherichi esherichi and G. esherichi elatior by the high prozona with distinct lobes and lateral projections of mesozona; to G. adaliae by the light, pale red (or pinkish-red) color of hind legs; to G. escherichi inermis and G. sevketi by the indistinct axillary 1 vein of male hind wing. But it can be separated from them by the following characters: It is different from G. escherichi escherichi and G. escherichi elatior by the shape of penis valves, the narrow and short metazona, shorter male tegmina, indistinct axillary 1 vein, the hind legs with light pale red color, small and slender body size and distribution. From G. escherichi inermis and G. adaliae by the shape of penis valves, the pronotum with distinct tubercles, high prozona with distinct lobes, distinct lateral projections of mesozona, narrow metazona with straight lateral margins and distribution. The other differences from G. adaliae by the narrower male tegmina, indistinct axillary 1; from G. escherichi inermis by the pinkish color of hind legs; differs from G. sevketi by the shape of penis valves, smaller pronotal tubercles, narrower metazona, light, pale red color of hind legs; smaller and slender body.

Distribution: ( Fig. 15). Turkey: Inner Aegean Region of Turkey: E of Manisa, SW of Kütahya, Uşak.

Remarks: The record given from Kütahya as “ escherichi ” ( Uvarov 1930) was noted by Cejchan (1965). He stated under G. escherichi escherichi that “On the basis of a single specimen it cannot be stated whether a separate taxonomic category or a mere individual variability” ( Cejchan 1965: 472). That specimen and one more male have the same label have been reidentified as this new taxon in the NHM. Those label data were written as “ Kütahya, Angora Prov. ”. Probably the word “ Angora ” (now Ankara) was erroneously added to the label. Kütahya is another province of Turkey quite far to Ankara and includes this new species.

The female paratype of G. escherichi elatior collected from Uşak ( Ramme 1951) is included to this taxon.

The specimens recorded as Eremobia Escherichi by Kuty (1907) were not examined. The collections from Kara-Tschehir (Karaşehir is in Tavşanlı: Kütahya) and Bos-Dagh (Bozdaǧ is in İzmir) may belong to this species rather than the others.

Etymology: Folk heroes, brave men who have lived in the Mountains of Aegean Region of Turkey called as “Efe” since 11th century. They mostly help people and save the Country against the enemies. They have particular folk dance known as “Zeybek” in Western Anatolia. The new species lives in the mountains of the Aegean Region of Turkey as an Efe.