Rhagonycha Eschscholtz, 1830

Kazantsev, S. V., 2023, A review of soldier beetles of the genus Rhagonycha Eschscholtz, 1833 (Coleoptera: Cantharidae) of Siberia, with description of a new species, Russian Entomological Journal 32 (3), pp. 271-296 : 288-294

Subgenus Rhagonycha Eschscholtz, 1830

Rhagonycha Eschscholtz, 1830: 64 .

type species: Cantharis fulva Scopoli, 1763 (designated by Delkeskamp, 1977)

= Nastonycha Motschulsky, 1853: 77 type species Nastonycha brachyptera

Motschulsky, 1853

= Pseudocratosilis Moscardini et Sassi, 1970: 192 type species Pygidia graeca Pic, 1901 (= Rhagonycha corcyrea Pic, 1901 )

= Spartiolepta Bedel et Bourgeois, 1901: 177 type species Telephorus geniculatus Lucas, 1846 (= Rhagonycha lucasi Jacobson, 1911 )

alpicola Barovskij, 1928: 98 . West and East Siberia (Altai, Tuva, Transbaikalia) – Middle Asia, Mongolia.

= kiritshenkoi Barovskij, 1929: 267

= shavrovi Barovskij, 1928: 99 Figs 92–97. Aedeagus of Rhagonycha : 92–94 — Rh. ussurica sp.n.; 95–97 — Rh. indistincta ; 92, 95 — dorsal view; 93, 96 — lateral view; 94, 97 — ventral view. Scale: 0.5 mm (95–97 — after Kazantsev, 1994).

Рис. 92–97. Эдеагус Rhagonycha : 92–94 — Rh. ussurica sp.n.; 95–97 — Rh. indistincta ; 92, 95 — сверху; 93, 96 — сбоку; 94, 97 — сниЗу. МасштабнаЯ линейка: 0,5 мм (95–97 — по: Kazantsev, 1994).

amaguensis Švihla, 1995: 74 . East Siberia (Irkutsk Oblast). = plumbea P. Rossi, 1792: 56

angulosa Kazantsev, 1994: 77 . East Siberia (Yakutia, = rhaetica Stierlin, 1862: 37

Transbaikalia). = sabauda Pic, 1914: 52

asiatica Wittmer, 1971: 195 . Far East (south of Primorskij = subrobusta Pic, 1923: 1

Krai) – Korea. atrovaria atrovaria Wittmer, 1971: 196 . West and East Siatra Linnaeus, 1767: 649 ( Cantharis ). Northwestern Sibe- beria (Altai, Tuva, Transbaikalia), Far East (Primorskij Krai) – ria (Khanty-Mansi A.O.) – centre and north of European part Kazakhstan, Mongolia. The only other subspecies is registered of Russia, Urals, Central Europe. in southern Urals (Mount Iremel)

= fauconetti Pic, 1914: 52 basarukini Kazantsev, 1994: 94 . Far East (Sakhalin) – Ja- = glacialis Bourgeois, 1909: 391 pan (Hokkaido).

Figs 98–103. Aedeagus of Rhagonycha : 98–100 — Rh. kabakovi ; 101–103 — Rh. cembricola ; 98, 101 — dorsal view; 99, 102 — lateral view; 100, 103 — ventral view. Scale: 0.5 mm (after Kazantsev, 1994).

Рис. 98–103. Эдеагус Rhagonycha : 98–100 — Rh. kabakovi ; 101–103 — Rh. cembricola ; 98, 101 — сверху; 99, 102 — сбоку; 100, 103 — сниЗу. МасштабнаЯ линейка: 0,5 мм (по: Kazantsev, 1994).

290 A review of Rhagonycha of Siberia cembricola Eschscholtz, 1822: 62 ( Cantharis ). East Siberia (Irkutsk, Chita Oblasts, Krasnoyarsk Krai, Yakutia), Far East (Kamchatka, Magadan Oblast, Khabarovsk and Primorskij Krais, Kuril Islands) – Mongolia, Korea.

= sjoestedti Pic, 1926: 2

coreana Pic, 1921: 28 . Far East (Primorskij Krai) – Japan, Korea dolini Švihla, 1995: 78 . East Siberia (Irkutsk Oblast), Far East (Sakhalin).

elongata Fallén, 1807: 11 ( Cantharis ). West and East Siberia (Khanty-Mansi A.O., Altai, Transbaikalia) – centre and north of European part of Russia, Urals, Western and Central Europe.

= atra Paykull, 1798: 266 ( Cantharis ) [homonym]

= paludosa L. Redtenbacher, 1847: 325 ( Cantharis ) [homonym]

flavotibialis Medvedev et Ryvkin, 1989: 138 . Far East (Amurskaya Oblast) – Mongolia.

fonticola Kazantsev, 1994: 83 . East Siberia (Evenkia, Yakutia), Far East (Magadan Oblast, Kamchatka, Amurskaya Oblast, Sakhalin, Kuril Islands ).

Figs 104–108. Aedeagus of Rhagonycha : 104–106 — Rh. nopporensis ; 107, 108 — Rh. basarukini ; 104, 107 — dorsal view; 105 — lateral view; 106, 108 — ventral view. Scale: 0.5 mm (after Kazantsev, 1994).

Рис. 104–108. Эдеагус Rhagonycha : 104–106 — Rh. nopporensis ; 107, 108 — Rh. basarukini ; 104, 107 — сверху; 99, 102 — сбоку; 100, 103 — сниЗу. МасштабнаЯ линейка: 0,5 мм (по: Kazantsev, 1994).

geniculata Gebler, 1832: 46 . East Siberia (Chita Ob- indistincta Medvedev et Ryvkin, 1989: 140 . East Silast), Far East (Kamchatka, Magadan, Amurskaya Oblasts, beria (Irkutsk, Chita Oblasts), Far East (Kamchatka, Primorskij Krai, Kuril Islands) – Japan (Hokkaido), Mon- Magadan, Amurskaya Oblasts, Primorskij Krai) – Mongogolia. lia, Korea.

= amurensis Pic, 1902: 25 kabakovi Kazantsev, 1994: 81 . Far East (Kuril Islands).

hyperborea Kazantsev, 1994: 84 . Far East (Magadan Ob- kurbatovi Kazantsev, 1994: 73 . Far East (south of Primorlast, Yakutia – Mongolia). skij Krai).

Figs 109–114. Aedeagus of Rhagonycha : 109–111 — Rh. mandibularis siberiana ; 112–114 — Rh. kurilica ; 109, 112 — dorsal view; 110, 113 — lateral view; 111, 114 — ventral view. Scale: 0.5 mm (after Kazantsev, 1994).

Рис. 109–114. Эдеагус Rhagonycha : 109–111 — Rh. mandibularis siberiana ; 112–114 — Rh. kurilica ; 109, 112 — сверху; 110, 113 — сбоку; 111, 114 — сниЗу. МасштабнаЯ линейка: 0,5 мм (по: Kazantsev, 1994).

292 A review of Rhagonycha of Siberia kurilica Wittmer, 1971: 197 . Far East (Kuril Islands) –

Northeastern China, Japan (Hokkaido). latiuscula J.R.Sahlberg, 1887:32 .West and East Siberia (Altai,

Kemerovo, Chita Oblasts, Krasnoyarsk Krai, Buryatia), Far East

(Magadan Oblast, Primorskij Krai, Sakhalin) – Japan, Mongolia. = caroli Pic, 1905: 121 = distinctipes Pic, 1905: 121 = mlikovskyi Švihla, 1995: 74 , syn.n. lederi Pic, 1909: 177 . Far East (‘Amur’, Khabarovsk Krai) –

North Korea. mandibularis siberiana Kazantsev, 1994: 93. East Siberia (Chita Oblast, Krasnoyarsk Krai), Far East (Kamchatka,

Magadan Oblast). The nominative subspecies, Rh. m. mandibularis (Kirby, 1837) , is distributed in North America, from Alaska and British Columbia to Newfoundland [ Delkeskamp, 1977; Kazantsev, 1994].

matisi Kazantsev, 1994: 85 . East Siberia (Chita Oblast, Yakutia), Far East (Magadan Oblast).

mimica Medvedev et Ryvkin, 1989: 140 . Far East (Amurskaya Oblast).

mongolica Wittmer, 1971: 197 . East Siberia (Tuva) – Mongolia.

nigriventris Motschulsky, 1860: 117 . West and East Siberia (Khanty-Mansi A.O., Tyumen Oblast, Taimyr, north of

Figs 115–120. Aedeagus of Rhagonycha : 115–117 — Rh. atrovaria ; 118–120 — Rh. hyperborea ; 115, 118 — dorsal view; 116, 119 — lateral view; 116, 119 — ventral view. Scale: 0.5 mm (after Kazantsev, 1994).

Рис. 115–120. Эдеагус Rhagonycha : 115–117 — Rh. atrovaria ; 118–120 — Rh. hyperborea ; 115, 118 — сверху; 116, 119 — сбоку; 111, 114 — сниЗу. МасштабнаЯ линейка: 0,5 мм (по: Kazantsev, 1994). Krasnoyarsk Krai, Buryatia, Yakutia), Far East (Kamchatka, = testacea Gyllenhal, 1808: 355 Magadan Oblast) – centre, north and south of European part of nopporensis Wittmer, 1971 c: 199 . Far East (Kuril Islands) Russia, northwestern Caucasus, Urals, Middle Asia, Western – Japan (Hokkaido). and Central Europe, Mongolia. pacifica Kazantsev, 1994: 80 . Far East (Sakhalin, Kuril Is- = femorata Rey, 1891: 115 lands) – Japan (Hokkaido). = innotatithorax Pic, 1902: 54 oboensis Kazantsev, 1994 b: 72 . East Siberia (Bolshie = limbata C.G. Thomson, 1864: 191 Koty, Irkutsk Oblast) – Mongolia. = nigrofemorata Schilsky, 1890: 178 planicollis Kazantsev, 1994: 76 . East Siberia (Chita Ob- = signicollis Rey, 1891: 115 last, Buryatia), Far East (Magadan Oblast, Kuril Islands).

Figs 121–126. Aedeagus of Rhagonycha : 121–123 — Rh. matisi ; 124–126 — Rh. transita ; 121, 124 — dorsal view; 122, 125 — lateral view; 123, 126 — ventral view. Scale: 0.5 mm (after Kazantsev, 1994).

Рис. 121–126. Эдеагус Rhagonycha : 121–123 — Rh. matisi ; 124–126 — Rh. transita ; 121, 124 — сверху; 122, 125 — сбоку; 123, 126 — сниЗу. МасштабнаЯ линейка: 0,5 мм (по: Kazantsev, 1994).

294 A review of Rhagonycha of Siberia selemdzhana Kazantsev, 2008: 337 . East Siberia (Amurskaya Oblast).

sibirica Wittmer, 1971 c: 198 . Far East (Primorskij Krai, Magadan Oblast) – Japan (Hokkaido).

sihotana Kazantsev, 1994: 75 . Far East (Primorskij Krai).

stusaki Švihla, 1992: 286 . East and West Siberia (Tyumen Oblast, Tuva, Buryatia, Yakutia) – northern Caucasus, Mongolia.

testacea Linnaeus, 1758: 403 ( Cantharis ). West and East Siberia (Khanty-Mansi A.O., Tomsk, Tyumen Oblasts, Krasnoyarsk Krai, Transbaikalia) – centre, north and south of European part of Russia, Urals, Western and Central Europe.

= florida Schiödte, 1872: 8

= obscurithorax Pic, 1902: 54

= ochropa Stephens, 1830: 296 ( Telephorus )

= pellucida Brahm, 1790: 110 ( Cantharis )

= taurinensis Pic, 1905: 105

transbaikalica Pic, 1906: 89 . West and East Siberia (Altai, Tuva, Yakutia, Transbaikalia) – Kazakhstan, Mongolia.

= jakovlevi Barovskij, 1908: 293

= smirnovi Barovskij, 1908: 294

transita Wittmer, 1971: 195 . East Siberia (Buryatia), Far East (Primorskij Krai, ‘Amur’) – Japan, Korea.

ulaensis Kazantsev, 1994: 78. East Siberia (Chita Oblast) – Japan (Hokkaido), Mongolia.

uralensis Dahlgren, 1972: 131 . East Siberia (Tuva).

ussurica sp.n. Far East (Primorskij Krai, Sakhalin).