Incasarcus viracocha Kury & Maury, 1998

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

3.35. Incasarcus viracocha Kury & Maury, 1998

Figs 4F , 9G-I , 18G-I , 30

Incasarcus viracocha Kury & Maury, 1998: 157 (desc.), 160 (key), figs. 38 (male dorsal habitus, chelicerae, pedipalpus, trochanter-patella IV), 39 (penis dorsal view), 40 (penis lateral view), 41 (male lateral habitus), 42 (female dorsal habitus), 43 (male femur IV); Kury 2003: 145 (cat.).

Redescription.

MALE: Measurements (n =3) DSW: 4.9-5.0 (4.9); DSL: 5.9-6.6 (5.9); CL: 2.4-3.0 (2.4). FIVL: 11.7-11.8 (11.8). ChL: 3.5-4.2 (4.2). Coloration (in ethanol): Brownish yellow body with reticular spots covering practically the entire carapace and chelicerae. Dorsum: (Fig. 4F ) Anterior margin of carapace with granular median elevation. Ocularium with very marked median depression; with sparse granules or smooth. Carapace with sparse granules. Areas I-IV with granules throughout their surface. Areas I-II and IV unarmed. Area III with a pair of median spines, facing posteriorly. Lateral and posterior margins of DS with granules throughout their length. Free tergites I-III with a row of granules. Chelicerae: (Fig. 4F ) Swollen in large males (as in the holotype). Segment I with densely distributed granules. Segment II with small granules or smooth; finger with five teeth. Segment III with three teeth. Pedipalpus : Covered with sparse granules across the dorsal surface of the femur-tibia. Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 6-7 small setiferous tubercles and a retro-subapical spine. Patella with a small retroapical setiferous tubercle. Tibia: prolateral iIiIIi, retrolateral IIii. Tarsus: prolateral iiiIiIIi, retrolateral iIiiIi. Venter: Coxa I with a median row of 5-6 setiferous tubercles. Coxae II-IV with setiferous granules. Rows of four tubercles between the coxae II-III and seven between the coxae III-IV. Genital area with few granules. Free sternites with transverse rows of small granules. Anal few granular. Legs: (Figs 4F , 9G-I ) Coxa I with a retrolateral and a prolateral apophysis. Coxa II with a prolateral apophysis. Coxa III unarmed. Coxa IV with sparse setiferous granules. Trochanters I-IV unarmed and granular. Femora I-III unarmed and with sparse granules. Femur IV with small granules; a retroventral row of 30-35 acuminate tubercles, distributed along the entire length of the article, except at the basal and apical ends (the most basal and apical tubercles are smaller than the median ones). Patellae I-III unarmed. Patella IV with a retroapical spiniform apophysis; a ventromedian row of four acuminate tubercles. Tibiae I-IV unarmed and granular. Tarsal formula: (n =3) 11, 21-22 (22), 11, 14-15 (15). Penis: (Fig. 18G-I ) VP rectangular, slightly tapering at the apex, with distal margin straight; straight in lateral view. MS C1-C3(C4) subapical long and apically curved; MS A1 median long and straight (⅓ of the length of the MS C); MS B1-B2 sub basal; MS B1 long and straight, MS B2 long and spatulated; MS D1 short, placed beneath MS C. Lateral sacs robust, with a blunt apex; with long T3-like microsetae. Stylus with slightly broad apex; with ventral projections. Promontory convex and tapered. - FEMALES: Measurements (n =3) DSW: 4.5-4.9; DSL: 5.5-5.8; CL: 2.0. FIVL: 12.4-12.5. ChL: 2.4-2.5. Chelicerae not swollen. Femur IV unarmed. Pedipalpal femur with five tubercles. Tarsal segmentation: (n =3) 9-10, 15-20, 10-12, 12-13.

Diagnosis.

It differs from other species of the genus by the set of following characters: DS without white spots on DS; DS granular; ocularium unarmed and with few granules or smooth; areas I, II and IV unarmed; area III with a pair of spines; free tergites I-III with a row of tubercles, unarmed (Fig. 4F ); male femur IV with a retroventral row of 30-35 acuminate tubercles (Fig. 9G-I ).

Distribution.

(Fig. 30 ) PERU. Cusco. Urubamba province.

Material examined.

Type material: Holotype ♂, ' PERU, Cusco, Urubamba province, Machu Picchu , 2,200-2,500m a.s.l., 13°07′S 72°34′W, 25/II/1994, J. Ochoa & J. Achicahuala leg. (MACN 9547) - Paratype ♀, ‘ditto’ (MACN 9548); ParatypeS 2 ♀, ‘ditto’ (MNRJ 5400). Additional material: 2 ♂, 3 ♀ ' PERU, Cusco’ (MUBI) .

Figures

Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury & Maury, 1998; C I. dianae Kury & Maury, 1998; D I. ochoai Kury & Maury, 1998; E I. pictus Kury & Maury, 1998; F I. viracocha Kury & Maury, 1998; Legend bars = 1 mm.

Figure 9. Male trochanter-patella / tibia IV of Incasarcus. A - B Trochanter-tibia IV of I. argenteus Kury & Maury, 1998, dorsal and ventral view, respectively; C - D Trochanter-tibia IV of I. dianae Kury & Maury, 1998, dorsal and ventral view, respectively; E-F Trochanter-tibia IV of I. pictus Kury & Maury, 1998, dorsal and ventral view, respectively; G - H Trochanter-tibia IV of I. viracocha Kury & Maury, 1998, dorsal and ventral view, respectively; I Trochanter-patella of I. viracocha, lateral view; J Trochanter-patella of I. ochoai Kury & Maury, 1998, large male, dorsal view; K - L Trochanter-tibia of I. ochoai, small male, dorsal and ventral view, respectively; M Trochanter-patella of I. ochoai, large male, lateral view; N Trochanter-patella of I. ochoai, small male, lateral view; Legend bars = 1 mm.

Figure 18. Penis of Huancabamba gen. nov. and Incasarcus, dorsal, lateral and ventral views, respectively. A - C H. kubricki gen. et sp. nov.; D - F I. ochoai Kury & Maury, 1998; G - I I. viracocha Kury & Maury, 1998; Legend bars = 0.1 mm.

Figure 30. Distribution of Incasarcus in Cusco, Peru.