Ummidia modesta (Banks, 1901)

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia modesta (Banks, 1901) Figs 27 , 28 , Map 4

Pachylomerus modestus Banks, 1901b: 570; HOLOTYPE: 1 ♂ (IZ22137) from Las Cruces, Dona Ana County, New Mexico, 32.338 -106.7661 5, 1238 m a.s.l. coll. Nathan Banks, deposited in MCZ, examined. PARATYPE: 1 ♀ (UMM586, AMNH) from Las Cruces, Dona Ana County, New Mexico, 32.338 -106.7661 5, 1238 m a.s.l., 23-27.ii.1937, coll. W.L. Chapel, AMNH.

Diagnosis.

Ummidia modesta can be differentiated from U. rosillos by the lack of a pale dorsal heart patch and brush on the retrolateral face of tarsus IV. Males can be differentiated from U. mercedesburnsae , U. paulacushingae , U. waunekaae , and U. gertschi by having a sinuous embolus and from U. paulacushingae , U. waunekaae , U. gertschi , U. timcotai , and U. rodeo by having relatively fewer spines on tibia I. Females can be differentiated from all other geographically proximate species except U. mercedesburnsae by having spermathecae with strong medial bend.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. In poor condition, specimen in many pieces. General coloration. Carapace and chelicerae reddish black 10R 2.5/1, legs dark reddish brown 5YR 3/4. Abdomen reddish black 2.5YR 2.5/1. Cephalothorax. Carapace 5.05 long, 4.79 wide. Pars cephalica 3.18 long. Foveal groove procurved, 0.33 long, 0.88 wide. Tubercle moderate. AER procurved. PER slightly procurved. Eye group 0.63 long, 1.12 wide, AME 0.26, PME 0.17, ALE 0.35, PLE 0.16. Sternum with marginal fringe, STRl 2.89, STRw 2.68. Chelicerae with anterior tooth row comprising five teeth, posterior margin with five teeth. Palpal endites with 13 cuspules spread over proximal half of endite face, lacking distal endite cuspules, ENDw 1.12, ENDl 2. Labium with five cuspules, LBw 0.91, LBl 0.77. Rastellum with many spines. Abdomen setose. Legs. F1 4.68; F1w 1.21; P1 2.14; Ti1 3.12; Mt1 2.05; Tr1 1.23; F3 3.47; F3w 1.41; P3 1.85; Ti3 1.94; Sd3 1.34; Mt3 2.07; Tr3 1.63; F4 4.62; F4w 1.32; P4 2.11; Ti4 2.96; Mt4 3.17; Tr4 1.59. Retrolateral face of tarsus IV with distinct comb of spinules which is surrounded in setae. Leg I spination pattern: TSp 7, TSpv 1, TSrd 0, TSr 0, TSrv 10, MtSp 2, MtSr 6, TrSp 3, TrSr 9. Pedipalps. PTl 3.03, PTw 1.04, Bl 2.58. Embolus relatively long, sinuous.

Description of female paratype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, and in vial with specimen. Specimen in poor condition, many limbs broken, in vial with specimen; right leg III missing. General coloration. Carapace, chelicerae, and legs dark reddish brown 2.5YR 2.5/4. Abdomen reddish black 10R 2.5/1, spinnerets strong brown 7.5YR 5/6. Cephalothorax. Carapace 9.86 long, 8.8 wide. Pars cephalica 6.74 long. Foveal groove procurved, 0.78 long, 2.17 wide. Eye tubercle moderate under median eyes. AER procurved PER straight. Eye group 1.47 long, 2.26 wide, AME 0.49, PME 0.38, ALE 0.6, PLE 0.42. Sternum sparsely setose around edges, thicker anteriorly, STRl 5.84, STRw 6.15. Chelicerae with anterior row comprising six teeth, posterior margin with five teeth. Palpal endites with 34 cuspules spread across proximal half of endite and 40 smaller cuspules distally, ENDw 2.25, ENDl 3.87. Labium with seven cuspules, LBw 2.3, LBl 1.6. Rastellum with many strong spines on process and up cheliceral face. Abdomen. Evenly setose. Legs. F1 6.14; F1w 2.26; P1 3.34; Ti1 3.94; Mt1 2.86; Tr1 1.4; F3 5.24; F3w 2.75; P3 3.49; Ti3 3.19, Sd3 2.08; Mt3 2.52; Tr3 2.36; F4 6.8; F4w 2.71; P4 3.9; Ti4 3.79; Mt4 4.07; Tr4 2.31. Retrolateral face tarsus IV with defined, tight comb with clusters of spinules and proximal and distal ends. Pedipalps. PF 5.44, PP 3.26, PTi 3.74, PTr 3.12. Spermathecae with medial bend, bulbs facing ventrally.

Material examined.

Known only from type material.

Figures

Map 4. Locality records for western United States and northern Mexico species.

Figure 27. Ummidia modesta (Banks, 1901) from New Mexico A male habitus illustration MCZ 22137 B female habitus illustration UMM 586. Scale bars: 4.0 mm.

Figure 28. Ummidia modesta (Banks, 1901) from Las Cruces, New Mexico. male holotype (MCZ 22137) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 586). Scale bars: 1.0 mm (A-D).