Spinulata, Davis & Gentili-Poole & Mitter, 2008

Davis, Steve R., Gentili-Poole, Patricia & Mitter, Charles, 2008, A revision of the Cossulinae of Costa Rica and cladistic analysis of the world species (Lepidoptera: Cossidae), Zoological Journal of the Linnean Society 154 (2), pp. 222-277 : 250-251

SPINULATA GEN. NOV.

Type species: Hemipecten acutipennis Schaus (1905) (by present designation).

Description

Male: Forewing length: 14–21 mm.

Female: Forewing length: 17–21 mm. Head: Antenna with inner serrate rami reduced in size; scales on antennal shaft cream to light brown; labial palpus cream to light brown ventrally, and light brown to brown dorsally; frons cream to brown, often cream and irrorated with rusty-brown; vertex cream to rusty-brown.

Thorax: Forewing often with fuscous spots at discal cell or along median of wing, and also often with obliquely transverse striae along entire wing or distal half of wing. R 2 of forewing most often fused and forked with R 3, sometimes separate; R 3 or R 2+3 of forewing always separate from R 4, never fused; R 4 of forewing most often separate from R 5, sometimes fused and forked.

Abdomen: Cream to rusty-brown dorsally; mostly cream to pale brown ventrally, but sometimes rusty-brown.

ering. Invagination between tergite 9 of genital capsule and uncus curving around base of gnathos. Gnathos with dorsum of bridge rugose; bridge broadly curved and arched as viewed posterioventrally; arms slightly protruding beyond bridge and mostly not recurving into tergite 9. Anal tube mostly membranous. Juxta process short, truncate and mostly fused to valva. Aedoeagus with dorsal hump present, but reduced in size; coecum reduced to nearly absent; rostellum digitiform and slender, with apex acute and rounded; vesica often covered with minute, setae-like spines.

Female genitalia: Sternite 8 variable, sometimes developed, sclerotized and triangular, with sides slightly concave, or sometimes reduced, mostly membranous, and covered by cuticular wrinkles of intersegmental membrane. Intersegmental membrane between abdominal sternites 7 and 8 often sclerotized and sculptured with many cuticular pits, wrinkles and folds. Ductus bursae slightly shortened to somewhat elongate. Corpus bursae reduced in size and oblong.

Etymology: Derived from the Latin ‘spinula’ meaning ‘thorn’. Spinulata refers to a patch of spines present terminally on the saccular process in the valvae of the male genitalia.

Male genitalia: Valva subtrapezoidal; saccular process a variably shaped spine patch, from triangular to an elongate, curving area of spines that stretch from the sacculus towards the costal area; base of costa with basal lobe strongly ascended dorsally, very pronounced and projecting at an angle. Uncus simple, often elongate, slender, and protruding beyond valvae, with a sparse setal cov- Discussion: Spinulata is proposed for four Costa Rican species that share two synapomorphies in the male genitalia. These are the broadly curved gnathos bridge, which is rugose in texture (character 23), and the development of a terminal spine patch on the saccular process of the valva (character 20). Superficially, Spinulata appears most allied to Simplicivalva , which differs in the almost complete absence of a saccular process.