Euboeus, Boieldieu, 1865

Nabozhenko, Maxim V., 2022, A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan, Zootaxa 5159 (4), pp. 451-486 : 482-483

Key to species of the genus Euboeus of the Caucasus, Iran and Turkmenistan, on males

1 Pronotum and head dark-bluish or dark-violet, elytra black ( Figs 19A, 20A, B)........................ E. chorasanicus

- Body dorsally completely black or sometimes black-brown................................................... 2

2 Puncturation of head and pronotum coarse and dense, punctures subequal or larger than interpuncture distance (sometimes punctures coarse but slightly sparser at middle of pronotum) ( Figs 1A, 2E, F, 4F, G, 5D, E, 6A, 7F, G). Male body dull dorsally ( Figs 1A, 2A, 3A, 4A, 5A, 7A) or rarely shiny ( Fig. 7A)...................................................... 3

- Puncturation of head and pronotum fine and sparse, puncture diameter much lesser than interpuncture distance, at least at middle and anterior third ( Figs 9D, E, 10A, B, 11E, F, 12D, E, 13E, F, 14D, E, 15D, E, 17D, E). Male body more or less shiny dorsally (8A, 9A, 10A, 11A, 12A, 13A, 14A, 15A, 17A, 18A)................................................. 7

3 Pronotum with very coarse and dense puncturation of longitudinal elongated punctures, especially on lateral sides of disc; Median smooth line of pronotum elevated near base and separated from other surface by impression with very dense merged punctures ( Figs 1A, 2E, F).................................................................. E. corrugosus

- Pronotum with coarse and dense puncturation of simple round, not merged punctures; median line not elevated near base or absent ( Figs 4F, G, 5D, E, 6A, 7F, G)..................................................................... 4

4 Head densely pubescent with recumbent reddish setae ( Figs 6A–D). Pro- and mesotarsi slightly widened, much narrower than apex of protibia ( Fig. 6A)....................................................................... E. vicinus

- Head bare, only with very short simple seta in each puncture ( Figs 4G, 5E, 7G). Pro- and mesotarsi stronger widened, equal or wider than width of apex of protibia ( Figs 4A, 5A, 7A)....................................................... 5

5 Metaventrite and abdominal ventrites pubescent with long recumbent hairs ( Fig. 4C, D). Pronotum very narrow P w P l = 1.13 ( Fig. 4F)................................................................................. E. gorganicus

- Metaventrite and abdominal ventrites bare, covered with simple very short setae ( Figs 5B, 7C, D). Pronotum wider, P w P l = 1.16–1.28 ( Figs 5D, 7F)............................................................................... 6

6 Pronotum much wider (P w P l = 1.28), with lateral strongly emarginated at basal quarter ( Fig. 5D). Abdominal ventrite 5 with double puncturation: large sparse punctures among fine dense puncturation ( Fig. 5B). Pro- and mesotarsi more elongate, pro- and mesotarsomere 1 longitudinal, pro- and mesotarsomere 2 with subequal length and width ( Fig. 5A)..... E. prometheus

- Pronotum narrower (P w P l = 1.16), with lateral margins straight or weakly rounded at base ( Fig. 7D). Abdominal ventrite 5 with simple fine and sparse puncturation ( Fig 7B). Pro- and mesotarsi wider, pro- and mesotarsomeres 1 and 2 transverse ( Fig. 7A).......................................................................................... E. zoroaster

7 Head pubescent with long recumbent setae ( Fig. 18E). Pronotum with projected, acute, pointed at apex antero-lateral corners ( Fig. 18D). Prothoracic hypomera with sparse distinct granules ( Fig. 18F)................................. E. zubovi

- Head bare ( Figs 8F, 9E, 11F, 12E, 13F, 14E, 15E, 17E). Antero-lateral angles of pronotum not projected, right or obtuse ( Figs 8E, 9D, 11E, 12D, 13E, 14D, 15D, 17D). Prothoracic hypomera with wrinkles and punctures, separately or together, but without sparse granules ( Figs 8G, 9F, 11G, 12F, 13G, 14F, 15F, 17F)................................................ 8

8 Lateral margins of pronotum not completely beaded, bead widely interrupted at middle ( Figs 17D, E). Apical piece of aedeagus subquadrate at apex or apex strongly emarginated at middle ( Fig. 17K)............................... E. quadricollis

- Lateral margins of pronotum completely beaded ( Figs 9E, F, 11E, F, 13E, F) or completely not beaded ( Figs 14D, E, 15D, E). Apical piece of aedeagus round at apex ( Figs 8D, 9K, 11M, 12K, 13M, 14K, 15K)................................. 9

9 Lateral margins of pronotum completely beaded ( Figs 9E, F, 11E, F, 13E, F)..................................... 10

- Lateral margins of pronotum not beaded ( Figs 8E, F, 12D, E, 14D, E, 15D, E).................................... 12

10 Pronotum widest ahead of middle ( Fig. 9D). Prothoracic hypomera not flattened along margin, with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background; wrinkles with microgranulation at anterior third ( Fig. 9F). Apical piece of aedeagus with longitudinal median keel ( Fig. 9K). Apical part of the basal piece of aedeagus smooth, without longitudinal rugosity ( Fig. 9K).............................................. E. grimmi

- Pronotum widest at middle ( Figs 11E, 13E). Prothoracic hypomera narrowly ( Fig. 13G) or widely ( Fig. 11G) flattened along margin, coarsely and densely punctured ( Fig. 13G) or coarsely longitudinally wrinkled ( Fig. 11G), without microgranulation. Apical piece of aedeagus with longitudinal median groove or wide impression ( Figs 11M, 13M)..................... 11

11 ody dorsally strongly shiny, especially pronotum with lacquer gloss ( Figs 10B, 11E, F). Prothoracic hypomera with coarse longitudinal wrinkles; lateral margin of prothoracic hypomera clearly widely flattened ( Fig. 11G). Metatibiae strongly bent ( Fig. 11C). Lateral margins of apical piece of aedeagus weakly rounded ( Fig. 11M)........................... E. huedepohli

- Body dorsally lesser shiny, pronotum without lacquer gloss ( Figs 13E, F). Prothoracic hypomera with coarse puncturation and smooth wrinkles at middle; lateral margin of prothoracic hypomera very narrowly weakly flattened ( Fig. 13G). Metatibiae very weakly bent ( Fig. 13B). Lateral margins of apical piece of aedeagus straight in apical half ( Fig. 13M)......... E. kasatkini

12 Prothoracic hypomera with large and coarse longitudinal smooth wrinkles ( Fig. 12F). Pronotum with strongly rounded lateral margins at middle ( Fig. 12D)................................................................. E. kalashiani

- Prothoracic hypomera punctured or with punctation and microwrinkles ( Figs 8G, 14F, 15F). Lateral margins of pronotum weakly rounded..................................................................................... 13

13 Prothoracic hypomera with dense uneven small uneven wrinkles and small sparse punctures on their background ( Fig. 8G)............................................................................................ E. arzanovi

- Prothoracic hypomera with large coarse foveolate punctures, without microwrinkles ( Figs 14F, 15F).................. 14

14 Eyes strongly convex (lateral margin angulate in dorsal view) ( Figs 14B, D). Pronotum wider (P w P l = 1.3), widest ahead of the middle, with moderately rounded lateral margins ( Fig. 14D). Base of flex side of meso- and metafemora with dense brush of setae ( Fig. 14B). Abdominal ventrites bare, with only simple very short setae ( Fig. 14B)................ E. krivokhatskyi

- Eyes moderately convex (lateral margin rounded in dorsal view) ( Figs 15B, D). Pronotum narrow, almost with subequal width and length (P w P l = 1.08), widest at middle, with very weakly rounded lateral margins ( Fig. 15D). Base of flex side of meso- and metafemora with simple recumbent setae, without separate dense brush ( Fig. 15B).......................... E. merkli