Hyrtanella Allen and Edmunds

Jacobus, Luke M. & Sartori, Michel, 2004, Review of the genus Hyrtanella (Ephemeroptera: Ephemerellidae), Zootaxa 785, pp. 1-12 : 2-3

Hyrtanella Allen and Edmunds

Diagnosis. Larvae are distinguished by having claws with basal denticles and a set of long subapical denticles; gills 3 operculate, with a median, transverse, weakened band; paired, blunt spines on abdominal terga 4–7; single, median spines on abdominal terga 8 and 9; and abdominal segments 8 and 9 reduced in width ( Allen & Edmunds 1976).

Female adults are distinguished by having a reduced number of marginal intercalaries between CuA and IMP in the forewing (one intercalary between IMP and MP2, and no intercalaries between MP2 and CuA), a relatively small hindwing with few crossveins and a sharp costal projection, a constricted abdomen, a medial spine vestige on abdominal segment 8, and enlarged abdominal segments 8 and 9 ( Allen & Edmunds 1976; Allen 1980).

Discussion. In the original description, Allen and Edmunds (1976) erroneously described gills present only on segments 3–6. As noted by McCafferty and Wang (2000) and Kluge (2004), Hyrtanella larvae indeed possess gills on segments 3–7.