Brachymesitius Barbosa, 2022

Barbosa, Diego N., Hermes, Marcel Gustavo & Lepeco, Anderson, 2022, Phylogeny of Mesitiinae (Hymenoptera: Bethylidae): assessing their classification, character evolution and diversification, Arthropod Systematics & amp; Phylogeny 80, pp. 603-625 : 603

Brachymesitius Barbosa gen. nov.

Type species.

Incertosulcus krombeini Móczár, 1970 by original designation.

Diagnosis.

The malar space convergent (#7:0), apex of median clypeal carina [in profile] inclined (#11:2), eye very small (#21:2), pubescence of eye absent (#22:0), posterior propodeal projection wide (#64:0), number of distal hamuli of hind wing four (#81:1), hypopygium wider than long (#94:2), anterolateral hypopygial apodeme present (#100:1) were found to be autapomorphies for Brachymesitius . The type species was first described as a species of Incertosulcus . The genus was characterized by the presence or absence of a long posterior propodeal projection, making the identification dubious. Azevedo et al. (2018) proposed a new interpretation for the diagnostic characteristics for Incertosulcus , making Incertosulcus krombeini , as junior synonym of Parvoculus indicus Kieffer, 1905.

This genus shares similarities with Anaylax , Clytrovorus , and Hadesmesitius which have the head, dorsal pronotal area and mesoscutum coriaceous, and the median pronotal line and median mesonotal sulcus absent. Other characteristics distinguish Brachymesitius from Anaylax , Clytrovorus , and Hadesmesitius , including eyes very small, similar to Bradepyris and Moczariella ; the anterolateral hypopygial apodeme similar to Mesitius ; and the hind wing with four distal hamuli is shared with Zimankos ; the presence of frontal carina and the propodeal declivity and lateral surface of metapectal-propodeal complex areolate are exclusive for Brachymesitius as diagnostic characteristics.

Description.

Wings: hyaline. Head: As long as wide; malar space as long as VOL, convergent; clypeus with median lobe rounded, median clypeal carina inclined; antenna with pubescence sparse and short; pedicel cylindrical, first flagellomere as long as pedicel, flagellomeres short; eye very small, without pubescence; frons foveolate, with frontal carina; ocelli very small; anterior ocellus crossing supra-ocular line; dorsal and ventral half of occipital carina low. Pronotum: Dorsal pronotal area shorter than wide, foveolate, with humeral angle rounded, side straight, anterior margin outcurved, posterior margin incurved, median pronotal line absent; mesoscutum coriaceous, median mesonotal sulcus absent, notaulus present and narrow; mesoscutellum not touching the metapectal-propodeal disc; metapectal-propodeal disc as long as its half width, metapostnotal median carina complete, with longitudinal ridge between metapostnotal median carina and metapostnotal-propodeal carina, posterior propodeal projection very short and thick; spiracle shape elliptical; propodeal declivity areolate, with median and lateral carinae; lateral surface of metapectal-propodeal complex areolate, without carinae. Wings: Hind wing with four distal hamuli. Metasoma dorsal and ventral region of terga III-VI polished; hypopygium bilobate, spiculum short, with lobate and short branch, wider than long, lateral margins convergent, with lateral anterior projection.

Etymology.

The name Brachymesitius , masculine, is a combination of the names “brachy”, from the Greek “short”, and refers to the reduced size of structures, such as eye size, flagellomeres, ocelli, length of dorsal pronotal area, posterior propodeal projection, and hypopygium, which are diagnostic for the group, and the name " Mesitius ", the type genus of Mesitiinae .

Distribution.

Iraq.

Species included.

Incertosulcus krombeini Móczár, 1970, now Brachymesitius krombeini ( Móczár, 1970) stat. rev. et comb. nov., removed from the synonym of Parvoculus indicus Kieffer, 1905.