Hyetussa Simon, 1902

Bustamante, Abel A. & Ruiz, Gustavo R. S., 2017, Systematics of Thiodinini (Araneae: Salticidae: Salticinae), with description of a new genus and twelve new species, Zootaxa 4362 (3), pp. 301-347 : 315-317

Hyetussa Simon, 1902

Hyetussa Simon, 1902: 363 (type species by original designation: Hyetussa cribrata Simon, 1902 = Hyetussa simoni Galiano, 1976 ); Galiano 1963b: 364 (redescription of type species); Galiano 1976a; Ruiz & Maddison 2015; Maddison 2015; World Spider Catalog 2017.

Mica Petrunkevitch, 1925: 246 [preoccupied by Mica Budde-Lund, 1908: 281 (Malacostraca: Isopoda); type species by original designation: Mica longithorax Petrunkevitch, 1925 ].

Micalula Strand, 1932: 141 (replacement name for Mica Petrunkevitch, 1925); Ruiz & Maddison 2015; Maddison 2015; World Spider Catalog 2017. NEW SYNONYMY.

Bredana Gertsch, 1936: 20 (type species by original designation: Bredana complicata Gertsch, 1936 ); Ruiz & Maddison 2015; Maddison 2015; World Spider Catalog 2017. NEW SYNONYMY.

Diagnosis. Males resemble those of Thiodina , Cyllodania , Arachnomura , Atomosphyrus and Tartamura in having spine-like setae on the anterior surface of male chelicera ( Figs 8A‒ 8I ; see also Galiano 1976b: figs 37, 39 and Galiano 1977a: fig. 11). They differ from Thiodina , Arachnomura , Atomosphyrus and Tartamura in having a complex embolus, coiled and lamellate at the tip ( Figs 16E‒16F , 17E‒17F ), and from Cyllodania in having a bump at the base (as in H. aguilari ) or in the middle of the embolus (see Figs 15C , 16D , 17D ; apical in Cyllodania ). Females resemble those of Cyllodania in the general pattern of genitalia, but differ in having more coiled copulatory ducts and smaller spermathecae ( Galiano 1976a: figs 12, 13).

List of species (chronological order):

1. Hyetussa cribrata (Simon, 1901) .

2. Hyetussa longithorax ( Petrunkevitch, 1925) comb. nov.

3. Hyetussa complicata ( Gertsch, 1936) comb. nov. 4. Hyetussa alternata ( Gertsch, 1936) comb. nov. 5. Hyetussa secta ( Mello-Leitão, 1944) .

6. Hyetussa simoni Galiano, 1976 (type species). 7. Hyetussa andalgalaensis Galiano, 1976 . 8. Hyetussa mesopotamica Galiano, 1976 . 9. Hyetussa aguilari Galiano, 1978 .

10. Hyetussa sergipe sp. nov.

11. Hyetussa tremembe sp. nov.

Note. The synonymy of Micalula and Bredana with Hyetussa is established by comparisons of the original descriptions ( Petrunkevitch 1925, Gertsch 1936), re-examination of the type species of Bredana , the neotype of Micalula longithorax and additional material with species currently placed in Hyetussa ( Galiano 1976a, 1976b, 1978). No substantial differences were observed to justify maintaining Micalula and Bredana as independent genera (see diagnosis of Hyetussa above).

Figures

FIGURE 8. Male chelicerae in Thiodinini, anterior view. A Arachnomura querandi sp. nov., holotype; B Cyllodania trinidad sp. nov., holotype; C Cyllodania zoobotanica sp. nov., holotype; D Hyetussa sergipe sp. nov., holotype; E Hyetussa tremembe sp. nov., holotype; F Tartamura agatelin gen. nov. et sp. nov., holotype; G Tartamura huao gen. nov. et sp. nov., holotype; H Tartamura metzneri gen. nov. et sp. nov., holotype; I Thiodina firme sp. nov., holotype; J Thiodina perian sp. nov., holotype; K Titanattus acanjuba sp. nov., holotype; L Titanattus euryphaessa sp. nov., holotype.Cyan arrows show spine-like setae.

FIGURE 15. Hyetussa complicata comb. nov. A male (FSCA), dorsal; B female (FSCA), dorsal; C male palp, slightly expanded, ventral view; D epigyne, ventral view.

FIGURE16. Hyetussasergipe sp.nov. A male holotype, dorsal; Bsame, lateral; C same, ventral;D male palp, ventral view; E same, retrolateral view; F same, retrodorsal view.

FIGURE17.Hyetussa tremembe sp. nov. A male holotype, dorsal;B same, lateral; C same, ventral; D male palp,ventral view; Esame, retrolateral view; F same, retrodorsal view.