Scipopus (Phaeopterina) heteropus ( Frey, 1927 ) Lindsay & Marshall, 2023

Lindsay, Kate & Marshall, Stephen A., 2023, A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae), European Journal of Taxonomy 904, pp. 1-189 : 130-131

Scipopus (Phaeopterina) heteropus ( Frey, 1927) comb. nov.

Fig. 57

Phaeopterina heteropus Frey, 1927: 74 .

Scipopus heteropus – Hennig 1934: 321 (key), 330 (listed).

Phaeopterinus heteropus – Aczél 1949: 342 (catalog). — Steyskal 1968: 48.15 (catalog). — Ferro & de Carvalho 2014: 60 (listed).

Differential diagnosis

Scipopus (Phaeopterina) heteropus appears to be identical to S. (Ph.) noturgidus sp. nov. in characters of the female terminalia but differs from S. (Ph.) noturgidus and other congeners by its black body and basally yellow mid and hind femora.

Type material examined

Syntypes

BOLIVIA • 1 ♂ (examined and photographed at NHMW 2002 ); NHMW • 1 ♀ (not examined); Yungas ; NHMW .

Other material examined

BOLIVIA • 2 ♀♀; La Paz , Chulumani, Apa Apa Reserve; 16°21′15″ S, 67°30′21″ W; 2000 m a.s.l.; 1 April 2001; S.A. Marshall leg.; DEBU (photographed, Fig. 57B–C, E ) • 1 ♀; same collection data as for preceding; DEBU0083 / DEBU083-06 sequenced for CO1; DEBU • 1?; Mapiri , Arroyo Tuhiri W; 15°29′08″ S, 68°25′8″ W; S.A. Marshall leg.; UGIC255-15/MYCRO156-15 sequenced for CO1–5; DEBU .

Description

LENGTH. 11–12 mm.

HEAD. Palpus light brown, entirely pale brown microtrichose and setulose, narrow (length 4.5× height). Clypeus dark brown or black, width ~2.2 × height, bare. Frontal vitta dull, orange, darker posteriorly, microtrichose. Epicephalon dark brown or black, shiny, bare, wide (width ⅔ or more of upper frontal vittal width at inner verticals), clearly delineated from upper frontal vitta. Postocellar seta absent, one lower fronto-orbital seta absent; all other head chaetotaxy well-developed.

THORAX. Scutum black, dark brown microtrichose. Female cervical sclerite slightly swollen anteriorly. Postpronotal lobe black, setulose on anterior half and outer lateral margin. Notopleuron black, with indistinct silvery microtrichose patch posterior to anterior notopleural seta. Thorax black, entirely silvery microtrichose. Legs black, yellow on basal ⅓ of mid femur, basal ½ of hind femur; fore tarsomere 1 black with prominent ventral golden fringe; hind tarsomere 1 black with slight anteroventral golden fringe. Wing evenly black-brown infuscate.

ABDOMEN (J+ ♀). T1 with long, fine black setae. Abdominal pleuron of available specimens discoloured.

FEMALE ABDOMEN. T1+2 ~1.8 × length of T3; posterior margin of T2 ~1.3× as wide as T1. Oviscape black, setulose, anterior ½ white microtrichose, ~3.5 × length of T6. Common duct absent, two paired spermathecal ducts arising separately from bursa copulatrix. Both paired spermathecal ducts short (3.0– 4.0× length of paired spermathecae), narrow, parallel-sided, swollen apically. Paired spermathecal stems narrow, longer than spermathecae, with sparse tubercles. Paired spermathecae approximately crescent-shaped, indented or pointed apically.

MALE ABDOMEN. T1+2 ~2.3 × length of T3. T6 short, genital fork 4.0 × as long as T6, with arms converging. Epandrium stout (length ≈ height); basiphallus, distiphallus, phallic bulb, phallapodeme not dissected (no male specimen available) and therefore not available for description.

Distribution

Brazil, Ecuador, Peru ( Steyskal 1968), Bolivia.

Figures

Fig. 57. Scipopus (Phaeopterina) heteropus (Frey, 1927). A. J, lateral view, Bolivia, syntype (NHMW). B. Head, ♀, lateral view, Bolivia, non-type (DEBU). C. Habitus, ♀, Bolivia, non-type (DEBU). D. Female spermathecae and associated structures (illustration credit: Sarah Schorno). E. Head, ♀, dorsal view, Bolivia, non-type (DEBU). F. Living, ♀, Bolivia. Abbreviations: pd = paired spermathecal duct; ps = paired spermatheca; pss = paired spermathecal stems.