Lodevia luganica ( Boyarina and Stschegolev, 1989 ) Boyarina, 1994

Boyarina, Natalya, 2010, Late Gzhelian pteridosperms with callipterid foliage of the Donets Basin, Ukraine, Acta Palaeontologica Polonica 55 (2), pp. 343-359 : 348-350

Lodevia luganica ( Boyarina and Stschegolev, 1989) Boyarina, 1994

Fig. 5C.

1989 Callipteris luganica sp. nov.; Boyarina and Stschegolev 1989: 101–104, pl. 2: a, b; text−fig. 2.

1994 Lodevia luganica ( Boyarina and Stschegolev, 1989) ; Boyarina 1994: 127.

Material.—Single fragment of the upper frond part— SNM PMC 2216/1; three pinnae fragments— SNM PMC 2216/122, SNM PMC 2216/123, and SNM PMC 2216/124.

Emended diagnosis.—Frond bipartite in the upper part, and bipinnate. Primary rachis bifurcating. Penultimate order axes longitudinally striated, finely tuberculate; overtopping. Pinnules palmate, ovate to oblong, decurrent. Pinnule margins crenulated, lobed to dissected into seven−nine segdoi:10.4202/app.2009.0020

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ments. Lobes or segments obovate, narrow, more or less arcuate. Venation pronounced. Midvein thick, terminating in the apical lobe or segment. Lateral veins making an acute angle to midvein, single or once bifurcating. Intercalated pinnules resembling the other pinnules; attached to the rachis between the pinnae.

Description.—The angle of the bifurcation is about 40 °. The penultimate order axes up to 2 mm wide. One completely preserved pinna of the penultimate order shows the overtopping of the axis and bears the stronger developed ultimate or− der pinnae on the external than on internal side. Pinnules vary in length from 8 to 24 mm. The pinnules, up to 8–10 mm long and 2–3 mm wide, are entire−margined to crenulated or lobed, strongly decurrent. The midvein is not very pronounced and bifurcates in the middle of the pinnule. The pinnules with a length of 12–24 mm and a width of 6–9 mm are dissected into seven−nine segments.

Stratigraphic and geographic range.—Upper Gzhelian (Upper Carboniferous) of Ukraine.