Amphineurus (Rhamphoneurus) sanus Alexander, 1929a

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.23. Amphineurus (Rhamphoneurus) sanus Alexander, 1929a

Amphineurus (Rhamphoneurus) sanus Alexander, 1929a: 189 (description, fig. 90: wing, fig. 225: male terminalia).

Material examined.

Holotype: 1 ♂, Chile, Chiloé Is., Ancud [41°52′S 73°48′W], 1927, F. & M. Edwards (BMNH). - Additional material: CHILE. 2 [sex unknown], Osorno , Pucotrihue [40°32′S 73°42′W], 10/ 12-IV-1968, Peña (USNM); 1 [sex unknown], Chiloé Is., Dalcahue [42°22′S 73°39′W], II-1954, Peña (USNM); 1 [sex unknown], Chiloé Is., Chepu [42°2′S 74°1′W], 4/ 6-IV-1968, Peña (USNM); 1 [sex unknown], Osorno , Bahia Mansa [40°35′S 73°44′W], South of Pucotrihue, 12-IV-1968, Peña (USNM); 1 ♂, Chiloé Is., Aulen [42°2′S 74°1′W], 8-II-1952, Peña (USNM) *.

Diagnosis.

This species is distinguished by a subtle spot on R2. Furthermore, this species has a remarkable sheath of aedeagus with a long horizontal spur, male tergite IX with a distinctive rectangular projection, mesal lobes absent and posterior margin of male tergite IX pigmented.

Redescription.

Wing length 5.86 mm, width 1.90 mm. - Coloration: General coloration dark brown. Rostrum and palpus dark brown. Antenna yellow, with apical segments slightly darker. Head brown. Eyes black. Thorax dark brown, pleura brown. Scutum without stripes. Halter brownish-yellow. Coxae brownish yellow and legs dark brown. Abdomen dark brown. - Head (dorsal view Fig. 28B , ventral view Fig. 28A ): Rostrum bifurcated with short appendices; first palpal segment shorter than terminal segment, terminal palpal segment dilated; scape narrower than pedicel; terminal flagellomere shorter and thicker than penultimate; antenna with several long setae. - Thorax (lateral view Fig. 28E , dorsal view Fig. 28F ): Anatergite taller than katatergite. Prothorax wide, prescutum with a few short setae. Halter with dilated knob. Wing (Fig. 28D ) almost clear of markings, except subtle marking on R2; veins around fork of bM (Fig. 28C ) faded; M1+2 longer than basal deflection of M1; R2+3+4 longer than R2+3. - Male terminalia (Fig. 28G ): Male tergite IX darkened on posterior margin, with wide median U-shaped notch; rectangular projection in two points on either side of extension; posterior margin with projection flat. Dorsal branch of gonocoxite very reduced. Lobe of gonostylus with pointed lobule shaped similar to folded leaf. Clasper of gonostylus with differently shaped branches: lateral branch long, bent around middle; medial branch curved into short stub. Mesal lobes absent. Sheath of aedeagus dark along almost entire length, angulated with smooth curvature, with setae on distal portion of lateral margin, as well as large distal horizontal lateral projection.

Remarks.

Some specimens were misidentified by C.P. Alexander as A. (R.) nullus Alexander. The rostrum of the holotype is longer than in the other species of this subgenus. This species resembles A. (R.) anchoralis sp. nov. but differs mainly in M1+2, shape of projection on male tergite IX, ventral branch of gonocoxite and spur of sheath of aedeagus.

Figures

Figure 28. Amphineurus (Rhamphoneurus) sanus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; sct, scutum; sctl, scutellum; t 9, male tergite IX.