Amphineurus (Rhamphoneurus) alexanderi, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.1. Amphineurus (Rhamphoneurus) alexanderi sp. nov.

Material examined.

Holotype: ♂, Chile, Llanquihue , Carelmapu [41°44′S 73°40′W], Coast of Canal de Chacao, 18-III-1957, Peña (USNM) *. Paratypes: 1 ♂, Chile, Chiloé Is., Dalcahue [42°22′S 73°39′W], 10/ 12-II-1954, Peña (USNM) *; 1 ♂, Chile, Chiloé Is., Aulen [42°2′S 74°1′W], 8-II-1952, Peña (USNM) *.

Etymology.

The species is named after C.P. Alexander, the remarkable Tipulomorpha expert who described all the species of this subgenus, among many others.

Diagnosis.

This species is distinguished by a curved R2+3, large markings only near R2, r-m longer than basal deflection of R5, and R2+3+4 longer than R2+3. Furthermore, the species has characteristic male terminalia with a trapezoidal-shaped spur and symmetrical tip of left mesal lobe.

Description.

Wing length 5.68 mm, width 1.61 mm. - Coloration: General coloration dark brown. Rostrum and palpus dark brown. First segment of antenna yellowish-brown, remaining segments dark brown. Head dark brown. Abdomen brown. - Head (dorsal view Fig. 8B , ventral view Fig. 8A ): terminal segments of antenna missing, as indicated by asterisks on Fig. 8B . Rostrum bifurcated with short appendices; first palpal segment shorter than terminal segment; scape shorter and thinner than pedicel. Thorax: Wing (Fig. 8C ) almost clear of markings, except strong marking covering R2; veins near fork of bM (Fig. 8D ) faded; r-m similar in length to basal deflection of R5; M1+2 shorter than basal deflection of M1; R2+3+4 longer than R2+3, more than twice length of R5. - Male terminalia (Fig. 8E ): Male tergite IX slightly darkened at posterior margin, with wide V-shaped median notch; posterolateral lobes triangular. Ventral branch of gonocoxite with one side with a few long setae, while other side with many short setae. Dorsal branch of gonocoxite half-length of ventral branch. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch wide club-shaped. Mesal lobes of gonocoxite asymmetrical; left mesal lobe with flat posterior margin; right mesal lobe elongated and curved. Sheath of aedeagus straight, darkened halfway of length; apex extended into narrowed tip and additional nearly rectangular preapical spur.

Remarks.

Specimens of this new species were previously identified by C.P. Alexander as A. (R.) insanus Alexander. This species resembles A. (R.) insanus but differs mainly in the attachment of m-cu, length of r-m, shape of thorax, mesal lobe and sheath of aedeagus.

Figures

Figure 8. Amphineurus (Rhamphoneurus) alexanderi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E Male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; cgonst, clasper of gonostylus; goncx, gonocoxite; l ms, left mesal lobe of gonocoxite; lgonst, lobe of gonostylus; r ms, right mesal lobe of gonocoxite; t 9, male tergite IX.