Stoebe phyllostachya (Candolle) Schultz Bipontinus (1861: 179)

Koekemoer, Marinda, 2016, A monograph of Stoebe and Seriphium (Gnaphalieae, Asteraceae) in southern Africa and neighbouring islands, Phytotaxa 242 (1), pp. 448-450 : 448-450

12. Stoebe phyllostachya (Candolle) Schultz Bipontinus (1861: 179) . Seriphium phyllostachyum Candolle (1838: 262) . Stoebe phleoides var. phyllostachya (Candolle) Harvey (1865: 282) . Type:— SOUTH AFRICA. C.B.S., Burchell 5131 (G-DC!, holotype; PRE!, K!, isotypes; P!, fragment).

= Seriphium phleoides Candolle (1838: 262) . Stoebe phleoides (Candolle) Schultz Bipontinus (1861: 179) . Type:— SOUTH AFRICA. C.B.S., Burchell 5052 (G-DC!, lectotype, designated here; K!, G!, BOL!, P!, isolectotypes). This specimen is a good representative of the species and has duplicates widely distributed.

= Stoebe copholepis Schultz Bipontinus (1861: 179) , syn. nov. Type :— SOUTH AFRICA. Caledon , Palmiet River, Ecklon 484 (56.3) (P!, lectotype, designated here).

= Stoebe ensori Compton (1934: 259) , syn. nov. Type:— SOUTH AFRICA. Cape Prov., Uniondale Div., Lauterwater (Long Kloof), dry hills, 750 m, 3 May 1933, Compton 4195 (BOL!, holotype; PRE!, K!, isotypes).

Multi-stemmed, woody shrubs, up to 0.4 m tall. Branches very densely or sparsely villous. Leaves linear-lanceolate, up to 10 mm long, mostly flat but also varying from almost flat to involute on the same plant, straight or slightly twisted; adpressed below the inflorescence, spreading lower down; abaxial surface with dense or sparse, long, flexible, adpressed hairs; adaxial surface hairs often geniculate, curled or with a loose loop in the middle; apices subaristate. Synflorescences globular to cylindrical, with interspersed subtending leaves, individual clusters of capitula and subtending leaves often clustered on the axis or much denser, and individual clusters not distinguishable. Capitula homogamous, discoid, 1-flowered. Receptacle reduced. Involucral bracts 8–14, in few rows, outer about half the length of the inner, leafy, sparsely or densely tomentose; innermost brown, scarious, acuminate to mucronate with a long aristate tip, the longest as long as or a little longer than the corolla tube. Florets bisexual, white or pink, tube 4 mm long; corolla lobes narrow, spreading, 1 mm long. Stamens 5, apical appendage lanceolate, basally tailed. Style bifid, branches not exceeding the corolla lobes, swollen at the base. Nectaries present. Cypselas short hairy, hairs adpressed, slightly longer towards the annulus. Pappus setae 12–15, slightly joined at the base, shorter than the corolla tube, plumose from close to the base, plumes fine, soft, ascending. Flowering March and April, occasionally in other months.

Distribution and habitat: —Southern Cape from Greyton to Kareedouw. Figure 30 .

Ecology: —Mountain slopes, at elevations of 100–1 000 m.

Conservation status: —Least Concern.

Notes: —Some specimens of this species are very similar to St. fusca , although St. fusca has yellow involucral bracts and does not have long, flexible hairs on the abaxial leaf surface.

Additional specimens examined:— SOUTH AFRICA, Western Cape. Cap, im Gebirge bei der Kapstadt (3318CD), Ecklon & Zeyher (84), (S); Montagu, Tradouws Pass, near toll house on Sleeping Beauty trail (3320DC), 26 January 1990, Koekemoer 338, (PRE); Swellendam div., Lemoenhoek Mountains, near Heidelberg (3320DD), March 1948, Stokoe SAM63076, (SAM); Riversdale, Garcias Pass, slope beyond Pass, Langeberg side (3321CC), 10 February 1982, Bohnen 8041, (NBG, PRE); Riversdale, Garcias Pass (3321CC), February 1904, Marloth 3571, (PRE); Oudtshoorn, Ruitersbos Forest Station, ridge between tributaries of Perdeberg River

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(3322CC), 18 March 1976, Hugo 114, (NBG, PRE); George, Outeniquas next to forestry track at Waboomskraal, 600m (3322CD), 1 March 1983, Vlok 546, (NBG, PRE); Uniondale, Many Waters Kloof (3323CA), 30 April 1935, Compton 5179, (NBG); Knysna dist., inter Knysna and Avontuur (3323CA/CC), 24 September 1930, Fries, Norlindh & Weimarck 1660, (K); Uniondale div, Long Kloof between Avontuur and sources of the Keurbooms River (3323CC), 18 March 1814, Burchell 5052, (BOL, G, K, P); George div., Keurbooms River, between Nuakamma River and Romanskraal (3323CC), 24 March 1814, Burchell 5131, (G-DC, K, P, PRE); Cloete’s Kraal, S of De Plugh (3323CC), April 1928, Fourcade 3763, (BOL); Tsitsikamma mountains, on southern slopes at Roode Muur (3323CC), 9 April 1989, Vlok 2106, (PRE); Knysna, next to Prince Alfred Pass, about 10 km south of Avontuur (3323CC), 6 March 1991, Vlok 2443, (PRE); George, Witte Els River (3323CD), 25 February 1944, Fourcade 6375, (BOL); Knysna, Deep River valley in dry macchia (3323CD), May 1924, Phillips 1078, (PRE); 2 km on Boesmanskloof road between Genadendal and Greyton (3419BA), 30 March 1996, Koekemoer 1269, (PRE); Bredasdorp, south of Napier (3419DB), 18 February 1951, Barker 7251, (NBG); Hills near Elim, 400ft (3419DB), July 1895, Bolus s.n., (BOL); Swellendam (3420AB), October 1897, Mund 107, (K, SAM); Swellendam (3420AB), October, Mund 366, (SAM); Swellendam, Zuurbraak (3420BA), January–March 1930, Thode A2337, (K, PRE); Knysna, Noetzie, false fynbos of coastal plateau, frequent on S aspect (3423AA), 14 September 1960, Acocks 21516, (K, PRE); Knysna, Concordia (3423AA), February 1922, Fourcade 2011, (BOL); Knysna (3423AA), January, Pappe SAM49693, (K, SAM); Plettenberg Bay, near Formosa Inn (3423AB), 2 April 1973, Botha 469, (PRE); Flats between Plettenberg Bay and The Poort (3423AB), April 1928, Fourcade 3739,

STOEBE AND SERIPHIUM ( GNAPHALIEAE )

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(BOL, NBG, PRE, S); Plettenberg Bay, E of airport, 60 m (3423AB), April 1976, Lavranos 12936, (E, PRE); Caledon, Palmiet River, Ecklon 484 (56.3), (P). Eastern Cape. Oudtshoorn, George, Kleinplaat-Duiwelskop area (3322DC), 15 February 1978, Bond 1356, (NBG, PRE); Uniondale, Die Hoek, N foot of Tsitsikama mts near Joubertina (3322DD), 9 March 1953, Esterhuysen 21247, (BOL); Uniondale, Die Hoek, N side of Zitzikama Mountains near Joubertina Valley and slopes, 2000–3000ft (3322DD), March 1954, Esterhuysen 22824, (BOL, K, PRE, S); Uniondale, Lauterwater, on dry hills (3323DC), 3 May 1933, Compton 4195, (BOL, K, PRE); Joubertina, Bruinklip, along foothills (3323DC), 15 July 1959, Van Breda 555, (BOL, K, PRE); Kareedouw Pass, S side (3324CD), January 1931, Fourcade 4554, (BOL, NBG);

Figures

FIGURE 30:—Known distribution of Stoebe phyllostachya in South Africa.