Stoebe leucocephala Candolle (1838: 259)

Koekemoer, Marinda, 2016, A monograph of Stoebe and Seriphium (Gnaphalieae, Asteraceae) in southern Africa and neighbouring islands, Phytotaxa 242 (1), pp. 448-450 : 448-450

7. Stoebe leucocephala Candolle (1838: 259) . Wigandia leucocephala (Candolle) Schultz Bipontinus (1861: 180) . Type:— SOUTH AFRICA. CBS, Rondeklip, Piekenierskloof n. Pretoriuskloof, 1500 ft, onder et Cederberg, 13 January 1831, Drège 5673 (G-DC!, lectotype, designated here, P!, isolectotype).

= Stoebe squarrosa Harvey (1865: 280) . Type :— SOUTH AFRICA. Lambert’s or Alexander’s Kloof, Wallich s.n. (TCD!, lectotype, designated here, K!, S!, isolectotypes).

Erect or diffuse shrubs or subshrubs, single or multi-stemmed, up to 1 m tall. Branches and leaves canescent. Leaves up to 15 mm long, variable: either narrow, linear, involute, twisted, curved inwards, often pigtail-like, or shorter and wider, with obtuse tips, twisted, ascending, both surfaces covered with adpressed hairs; woolly hairy when young; abaxial surface becoming glabrous with age; pleasantly aromatic. Synflorescences of many capitula very tightly massed in round, terminal heads. Capitula homogamous, discoid, 1-flowered. Receptacle reduced to short stalks on a swollen globose base. Involucral bracts white, opaque, 12; outer 3 bracts short, about half the length of the inner bracts, upper part leafy; inner bracts milky white, up to 6.0 × 0.7 mm, narrowly lanceolate; tips rounded to acute. Florets bisexual, corolla a deep pink to plum-red; corolla lobes narrow, erect, less than one third the length of the tube, inner margins with a row of distinct, large, inflated cells. Stamens 5; apical appendage lanceolate, basally tailed. Style bifid, somewhat swollen at the base. Nectaries present. Cypselas white, silky-hairy, hairs very thin, long, flexible, adpressed, twisted, tips very fine, acuminate; annulus a distinct rim of cypsela hairs. Pappus setae 20–24, slightly fused at the base; plumose from the base; plumes adpressed, very short at the base becoming progressively longer upwards, slightly tufted at the tips. Flowering November and December, occasionally in October and January.

Distribution and habitat: — Western Cape from Nieuwoudtville to Malmesbury, West of the meridian 20?. Figure 25 .

Ecology: —In deep sand and sandstone hills, at elevations of 200–800 m.

Conservation status: —Least Concern.

Additional specimens examined:— SOUTH AFRICA, Western Cape. Van Rhynsdorp Div., Klaver (3118DC), 29 October 1944, Leipoldt 4130 (PRE); Van Rhynsdorp div., Klaver (3118DC), 29 October 1944, Leipoldt 4131 (BOL); Van Rhynsdorp, Gifberg near the top of Ou Berg Pass (3118DD), 20 November 1995, Koekemoer 1103 (PRE); Clanwilliam, on plateau on summit of Nardouw Mountains (3118DD), 29 November 1910, Pearson 5423 (BOL, K, NBG); Lokenburg, arid fynbos on TMS ridges W of river, sandy valley (3119CA), 12 October 1953, Acocks 17357 (PRE); Calvinia dist., Lokenburg, 21 miles S of Nieuwoudtville (3119CA), 12 October 1953, Story 4331 (PRE); Top of Nardouwsberg (3218BB), 20 March 1996, Koekemoer 1218 (PRE); Onder Bokkeveld und Cederberg (3218DB), 1835, Drege 5673 (PRE); Piekenierskloof (3218DB), Drege SAM16822 (SAM); Clanwilliam, Heuningvlei—Pakhuis, N Cederberg (3219AA), 16 January 1953, Esterhuysen 21145 (BOL, K, PRE); Clanwilliam div., Pakhuis Mts (3219AA), 28 December 1941, Esterhuysen 7403 (BOL, K); Clanwilliam, Cederberg, Heuningvlei (3219AA), December 1941, Stokoe SAM58124 (SAM); Heuningvlei, roadside above locked Forestry gate at boundary, ca 25 m above gate (3219AA), 22 October 1985, Taylor 11253 (NBG, PRE); 24 km from Clanwilliam on Cederberg Rd. (3219AC), 21 November 1995, Koekemoer 1107 (PRE); Clanwilliam div.,

66 • Phytotaxa 242 (1) © 2016 Magnolia Press

KOEKEMOER

Elandskloof W end (3219CA), 20 December 1940, Esterhuysen 4206 (BOL); Clanwilliam, Elandskloof (3219CA), December 1940, Stokoe SAM58123 (SAM); Malmesbury, Rondevlei, W of Preekstoel (3318BC), 2 December 1977, Esterhuysen 34790a (BOL); La Fonteine, a farm east of Riebeeck Kasteel town, local patch of coastal fynbos (3318BD), 5 November 1976, Boucher 3253 (NBG, PRE); Malmesbury dist, in fynbos stand E of Riebeek Kasteel Village (3318BD), 25 November 1976, Esterhuysen 34479 (BOL, S); Malmesbury, Burgerspost farm, Pella Fynbos Biome site near plantation along N-E boundary (3318DA), 23 December 1981, Boucher 5048 (NBG, PRE); Malmesbury div., 1.5 miles west of Kalabas Kraal (3318DA), 8 December 1934, Salter 5100 (BOL, K, SAM); Calvinia div., Uien Valley, Bokkeveld Mts., 2000–2500ft, December, Drege (K); Uienvalei, in der Valei und auf felsiger Höhe, 5 December 1839, Drege (P, S); Cap, Drege (G); Prom. B. Spei, 1839, Drege (G); Port Natal et Afr. Meridionale, Drege (G); CBS, Drege (E); Rondklip; Piekenierskloof n. Pretoriuskloof, 13 January 1831, Drege 5673 (P); Little Namaqualand, roadside Middelkraal, 7 December 1910, Pillans 5606 (K); CBS, Wallich (K, S, TCD).

Figures

FIGURE 25:—Known distribution of Stoebe leucocephala in South Africa.