Euphoria fulveola Bates, 1889

Orozco, Jesús, 2012, Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae), The Coleopterists Bulletin (mo 11) 66, pp. 1-182 : 64-65

Euphoria fulveola Bates, 1889

(Appendix 4: Fig. 30 )

Euphoria fulveola Bates 1889: 370 . Original combination.

Lectotype at BMNH designated by Hardy (2001), examined. Two paralectotypes at BMNH, not examined.

Description (n = 60). Length 10.5–12.9 mm; width 6.0–8.0 mm. Color: Dorsal surface yellowish brown, tomentous, rarely shiny. Pronotum with 1 dark vitta at each side of midline; vittae longitudinal, black, covering most or all of surface. Elytra with black irregular markings, occasionally covering elytra almost entirely. Ventral surface, including legs, light brown to black, abdominal sternites sometimes bicolored. Head: Frons frequently slightly raised at middle, densely punctate; punctures round, deep, moderate in size, confluent, moderately densely to densely setose; setae, yellowish, moderate to long. Clypeus subrectangular, lateral declivity moderately to strongly expanded, sides strongly raised in males, weakly raised or flat in females; apex truncate in dorsal view, sinuate in frontal view, weakly reflexed in males, vaguely reflexed in females. Antennal club longer than stem in males, shorter in females. Pronotum: Surface moderately densely to densely punctate; punctures round to lunulate, small to moderate, sparsely to densely setose; setae short to moderate, yellowish. Sides at base subparallel to moderately convergent anteriorly, anterior corners with few rugae. Base in front of scutellum weakly to strongly emarginate. Scutellum sparsely to moderately punctate; punctures small, lunulate, frequently bearing setae. Elytra: Surface moderately densely punctate, striae bearing 2 densely punctate,lateral rows,and 1 sparsely punctate, medial row, punctures small, lunulate to irregular, glabrous to sparsely setose; setae minute to short, whitish to yellowish; apex weakly rugose. Pygidium: Surface subconcentrically striate; striae dense, discontinuous, sparsely to moderately densely setose, setae as on pronotum. Legs: Protibiae slender in males, robust in females, teeth frequently equidistant, basal tooth frequently weakly developed in males. Mesotibial carina moderately to strongly developed, often ending in 1 or 2 sharp spines. Metatibiae not apically expanded in males, strongly expanded in females; metatarsi longer than metatibiae in males, as long or shorter in females; spurs slender in males, at least 1 thick or expanded in females. Venter: Mesometasternal process moderately compressed laterally, extending anteriorly slightly or well beyond mesocoxae, apex variably rounded to truncate. Mesepimera, metasternum, and metacoxae setose, setae as on legs. Metasternum rugose, moderately to densely setose laterally, weakly to moderately punctate and setose at middle; median sulcus strongly impressed. Abdomen in lateral view flat to vaguely concave in males, weakly to moderately convex in females. Male genitalia: Parameres as in Fig. 30c .

Diagnosis. This species is separated from other species in the group by the weakly raised clypeal apex in males, elytral striae bearing two densely punctate lateral rows and one sparsely punctate medial row composed of small lunulate and irregular punctures, and form of the parameres. Based on the material examined, females of this species cannot be separated morphologically from females of E. avita . Nevertheless, the two species are known from different geographic areas.

After E. vestita , this species was found to be the most frequently given in error in the literature and on identification labels to several species in the avita and geminata species-groups.

Taxonomic History. Bates (1889) described the species based on seven specimens from “ Mexico, Zacultipan in Hidalgo, Oaxaca ” and commented on the similarities with E. avita and E. vestita . Hardy (2001) erroneously listed Euphoria nigriventris Bates , a synonym of Euphoria humilis Blanchard , as a synonym of this species.

Natural History. Adults have been collected in pine-oak forests and attracted to fruit traps. The species is known to occur between 1,160–2,228 m elevation.

Temporal Distribution. June (36), July (12), August (1), September (2) ( Fig. 30e ).

Geographic Distribution. Known from Hidalgo, Oaxaca, Puebla, Querétaro, and San Luis Potosí in Mexico ( Fig. 30f ).

Specimens Examined (60). Type material: Euphoria fulveola Bates, 1886 ; lectotype at BMNH labeled “LECTO-/TYPE// SYN-TYPE// Oaxaca / Mexico / Hoege// Euphoria / fulveola/ Bates// B.C.A. Col., II(2)./ Euphoria // LECTOTYPE / Euphoria / fulveola/ Bates = By / A.R. Hardy , 78” . Other material: MEXICO (59): HIDALGO: Cardonal (1), Chapulhuacán (1), Jacala (18), Maguey Verde (3), Molango (2), Santuario (15), Tulancingo (1), Zacualtipán (2), Zimapán (1); OAXACA: Oaxaca (1); PUEBLA: La Cumbre (2), Xicotepec de Juárez (2); QUERÉTARO: El Madroño (1), Jalpán de Serra (1); SAN LUIS POTOSÍ: San Luis Potosí (1), Xilitla (7).

Figures

Fig. 30. Euphoria fulveola. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in Mexico.