Cladiopsocus, Roesler, 1940

Obando, Ranulfo González, García Aldrete, Alfonso N. & Carrejo, Nancy, 2018, New species and records of Cladiopsocus Roesler (Psocodea: ‘ Psocoptera’: Psocomorpha: Cladiopsocidae) from Colombia, Zootaxa 4378 (2), pp. 191-223 : 192

publication ID	https://doi.org/ 10.11646/zootaxa.4378.2.2
publication LSID	lsid:zoobank.org:pub:0FAAA8FA-83EC-404E-9C44-6ED1355B7FF7
DOI	https://doi.org/10.5281/zenodo.5959330
persistent identifier	https://treatment.plazi.org/id/F41EAC0D-FFCB-0A1E-0F9F-FEDDEAB2F856
treatment provided by	Plazi
scientific name	Cladiopsocus
status

Treatment

Key to neotropical species of Cladiopsocus View in CoL (males)

(Modified from González et al. 2016)

Males unknown of C. clarus , C. dolosus , C. fuscus , C. nebulosus and C. ocotensis

1. Paraproctal tubercle enlarged into a distinct “thumb” ( Figs 21 View FIGURES 15–21 , 49 View FIGURES 43–49 , 89 View FIGURES 83–89 ) or enlarged but without a distinct “thumb” ( Figs 56 View FIGURES 50–56 , 82 View FIGURES 76–82 ).................................................................................................... 2

- Paraproctal tubercle reduced, not enlarged nor with a distinct “thumb” ( Figs 7 View FIGURES 1–7 , 14 View FIGURES8–14 , 28 View FIGURES 22–28 , 35 View FIGURES29–35 )........................... 19

2. Forewing veins with spurs ( Figs 15 View FIGURES 15–21 , 57 View FIGURES 57–62 , 76 View FIGURES 76–82 ), sometimes restricted only to the margin ( Figs 50 View FIGURES 50–56 , 76 View FIGURES 76–82 ); epiproct emarginated apically ( Fig. 20 View FIGURES 15–21 ), or with a broad posterior process ( Figs 55 View FIGURES 50–56 , 81 View FIGURES 76–82 ).................................................. 3

- Forewing veins without spurs ( Figs 43 View FIGURES 43–49 , 83 View FIGURES 83–89 ); epiproct emarginated apically ( Fig. 48 View FIGURES 43–49 ) or with bumps, prongs, papillae or flanges................................................................................................... 12

3. Paraproctal tubercle clearly enlarged into a distinct “thumb” ( Figs 21 View FIGURES 15–21 , 49 View FIGURES 43–49 , 89 View FIGURES 83–89 ); epiproct emarginated posteriorly ( Figs 20 View FIGURES 15–21 , 5 View FIGURES 1–7 0, 88), sometimes with a broad posterior process ( Figs 55 View FIGURES 50–56 , 81 View FIGURES 76–82 ).................................................... 4

- Paraproctal tubercle enlarged but without a distinct “thumb” ( Figs 56 View FIGURES 50–56 , 82 View FIGURES 76–82 ); epiproct emarginated or with a broad posterior process ( Figs 55 View FIGURES 50–56 , 81 View FIGURES 76–82 )...................................................................................... 8

4. Paraproctal prong proximally with a denticulate process on the inner margin; epiproct emarginate posteriorly ( Fig. 21 View FIGURES 15–21 ).......................................................................................... C. caguanensis n. sp.

- Paraproctal prong proximally without a denticulate process on the inner margin ( Figs 132, 134 View FIGURES 120–136 )....................... 5

5. Paraproctal prong distally forked, with arms pointed ( Fig. 132 View FIGURES 120–136 ); epiproct obtusely concave posteriorly, postero-lateral corners pointed, setose ( Fig. 105 View FIGURES 103–119 )........................................................... C. panchei González et al.

- Paraproctal prong not distally forked, or with a short preapical tooth ( Fig. 134 View FIGURES 120–136 ); epiproct variable, with or without a distal process ( Fig. 116 View FIGURES 103–119 )........................................................................................ 6

6. Paraproctal prong with a short preapical tooth ( Fig. 134 View FIGURES 120–136 ); epiproct trapeziform, with widely emarginated posterior margin ( Fig. 116 View FIGURES 103–119 )........................................................................... C. julianae González et al.

- Paraproctal prong without a preapical tooth ( Figs 126, 130 View FIGURES 120–136 ), epiproct variable, sometimes trapeziform ( Fig. 103 View FIGURES 103–119 ), semi-oval, emarginated posteriorly or modified ( Fig. 118 View FIGURES 103–119 ).............................................................. 7

7. Epiproct trapeziform, posterior border projected, slightly concave ( Fig. 103 View FIGURES 103–119 ); paraproctal prong with a distal tooth, and a small proximal denticle on the inner margin ( Fig. 126 View FIGURES 120–136 )...................................... C. arboricolus Calderón et al.

- Epiproct strongly bilobed and raised distally ( Fig. 118 View FIGURES 103–119 ); paraproctal prong without a small proximal denticle on the inner margin ( Fig. 130 View FIGURES 120–136 )...................................................................... C. ramulosus (Enderlein)

8. Forewings with well developed spur veins (see Fig. 55 View FIGURES 50–56 in González et al. 2016); paraproctal prong single, pointed ( Fig. 133 View FIGURES 120–136 ); epiproct projected posteriorly, emarginate, postero-lateral corners rounded ( Fig. 115 View FIGURES 103–119 )............ C. tikunus González et al.

- Forewings with spur veins poorly developed ( Fig. 15 View FIGURES 15–21 ); paraproctal prong single or double ( Fig. 19 View FIGURES 15–21 ); epiproct variable...... 9

9. Paraproctal prong double ( Fig. 82 View FIGURES 76–82 ); epiproct with widened median posterior projection and rounded or emarginate posterior margin ( Fig. 81 View FIGURES 76–82 ); phallosome well sclerotized, with internal parameres as two narrow, sclerotized arches, touching each other on the midline ( Fig. 80 View FIGURES 76–82 )................................................................................ 10

- Paraproctal prong single ( Fig. 56 View FIGURES 50–56 ) or with rugose processes, epiproct emarginated or with posterior median process rounded distally; phallosome variable............................................................................ 11

10. Paraproctal prong with inner arm short, truncate ( Fig. 129 View FIGURES 120–136 ); epiproct with median process slightly emarginate posteriorly, postero-lateral corners round, small, slightly projected ( Fig. 108 View FIGURES 103–119 ); phallosome with internal parameres as two narrow arches (see Fig. 21 View FIGURES 15–21 in González et al. 2016).................................................... C. eertmoedi González et al.

- Paraproctal prong with inner arm almost as long as the outer arm, tapered ( Fig. 82 View FIGURES 76–82 ); epiproct with posteriorly rounded median process ( Fig. 81 View FIGURES 76–82 ); phallosome with internal parameres narrower than in the previous species ( Fig. 80 View FIGURES 76–82 )........ C. sixtoi n. sp.

11. Epiproct emarginated posteriorly; paraproctal prong with rugose processes......................... C. uncinatus (New)

- Epiproct with posterior median process of rounded apex ( Fig. 55 View FIGURES 50–56 ); paraproctal prong long, curved and tapered distally ( Fig. 56 View FIGURES 50–56 )............................................................................... C. norsantanderinus n. sp.

12. Paraproctal prong with a laminar oval process between it and the mesal tubercle ( Figs 89 View FIGURES 83–89 , 135 View FIGURES 120–136 ). Epiproct posteriorly emarginated, postero-lateral corners widely rounded ( Figs 88 View FIGURES 83–89 , 119 View FIGURES 103–119 ).................................................... 13

- Paraproctal prong without oval process between it and the mesal tubercle ( Fig. 49 View FIGURES 43–49 ). Epiproct variable in shape ( Fig. 48 View FIGURES 43–49 )... 14

13. Paraproctal prong short, stout, distally blunt ( Fig. 135 View FIGURES 120–136 )............................. C. amacayacuensis González et al.

- Paraproctal prong long, distally pointed ( Fig. 89 View FIGURES 83–89 )................................................ C. tinigua n. sp.

14. Paraproctal prong short, widened distally, with two-three apical teeth ( Fig. 49 View FIGURES 43–49 ), epiproct wider than long, posterior margin emarginated ( Fig. 48 View FIGURES 43–49 ).................................................................. C. laesperanza n. sp.

- Paraproctal prong well developed, pointed or stout and with one-three apical teeth; epiproct variable in shape........... 15

15. Epiproct with small posterior process ( Fig. 109 View FIGURES 103–119 ); paraproctal prong curved, with one apical tooth ( Fig. 125 View FIGURES 120–136 ).............................................................................................. C. pacificus Calderón et al.