FISSURELLIDAE J. Fleming, 1822

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 580-581

Family FISSURELLIDAE J. Fleming, 1822

auversiensis , Emarginula – Deshayes, 1861-DescrCoqFoss: 248; 1862-DescrCoqFoss: 27, as Emarginula “ caelata ”, pl. 27, figs 1-4 [given in text as pl. 22, fig. 1]; 1865-DescrCoqFoss: 665 [error on p. 248 corrected]. — Type locality: Auvers-sur-Oise. — Type age: Eocene (Bartonian). — Current status: Emarginula auversiensis Deshayes, 1861 ( Le Renard & Pacaud 1995: 86).

brevirimata , Emarginula – Deshayes, 1863: 46-47, pl. 6, figs 16a, 17-18. — Type locality: La Réunion. — Type age: Recent. — Type material: MNHN-IM-2000-4744, holotype. — Current status: Montfortula brevirimata (Deshayes, 1863) ( Herbert 1987: 4-6, figs 9-12; McLean 2011: 417).

caelata , Emarginula ; see: auversiensis , Emarginula clathrata , Emarginula – Deshayes, 1824-DescrCoqFoss: 17-18, 1, pl. 1, figs 26-28;1830b-EncyMeth: 111-112; 1836-HistNatAnim2: 588-589; 1843-HistNatAnim3: 217; 1861-DescrCoqFoss: 249. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Current status: Emarginula clathrata Deshayes, 1824 ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 41).

costaria , Fissurella – Deshayes, 1824-DescrCoqFoss: 20, 2, pl. 2, figs 10-12; 1824-DictClass6: 527; 1830b-EncyMeth: 135; 1833b: 232; 1835b: 134; 1836-HistNatAnim2: 602-603; 1843-HistNatAnim3: 221-222; 1850-TraitElem: 38, pl. 64, figs 7-9. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). This species is not present in the Eocene of Paris Basin. — Current status: synonym of the Miocene to Recent Diodora italica (Defrance, 1820) [ Fissurella ] ( Sabelli et al. 1990: 124).

costulata , Emarginula – Deshayes, 1863: 47, pl. 7, figs 3-5. — Type locality: La Réunion. — Type age: Recent. — Type material: MNHN-IM-2000-4750, lectotype;MNHN-IM-2000-4751, paralectotype. — Current status: Emarginula costulata Deshayes, 1863 ( Herbert 1987: 11-12, figs 39-42; Herbert 2015: 21).

curvirostris , Emarginula –Deshayes, 1830b-EncyMeth: 111; 1835b: 134; 1836-HistNatAnim2: 586;1843-HistNatAnim3: 216. — Type locality: Mediterranean. — Type age: Recent. — Type material: MNHN-IM-2000-4752, 3 syntypes. — Current status: synonym of Emarginula sicula J. E. Gray, 1825 ( Sabelli et al. 1990: 125).

cymbiola , Emarginula – Deshayes, 1861-DescrCoqFoss: 249, 4, pl. 4, figs 5-8. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Current status: Entomella (Entomella) cymbiola (Deshayes, 1861) ( Le Renard & Pacaud 1995: 87) . Entomella cymbiola (Deshayes, 1861) ( Pacaud 2004: 607; Pacaud 2008: 41; Pacaud 2017c: 183-184).

decisa , Fissurella –Deshayes, 1861-DescrCoqFoss: 238-239, 7, pl. 7, figs 13-16. — Type localities: Auvers-sur-Oise, Le Fayel, Boucouvilliers & Le Guépelle. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32390, syntype from Auvers-sur-Oise. — Current status: Diodora decisa (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86; Pacaud 2017c: 182-183).

decorata , Emarginula – Deshayes, 1863: 47-48, pl. 7, figs 6-8. — Type locality: La Réunion. — Type age: Recent. — Type material: MNHN-IM-2000-4754, holotype. — Current status: Emarginula decorata Deshayes, 1863 (Manousis & Galinou-Mitsoudi 2014: 3, 10, fig. 2a, as being introduced into the eastern Mediterranean ; Herbert 2015: 21, fig. 3E-G) .

defrancii , Rimula – Deshayes, 1861-DescrCoqFoss: 244, 3, pl. 3, figs 233-236. — Type locality: Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Current status: Rimula defrancii Deshayes, 1861 ( Pacaud 2008: 42).

denudata , Fissurella – Deshayes, 1861-DescrCoqFoss: 241-242, 7, pl. 7, figs 1-4. — Type localities: Parnes & Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Current status: Diodora denudata (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 42).

distans , Fissurella – Deshayes, 1861-DescrCoqFoss: 240, 7, pl. 7, figs 28-30. — Type localities: Cuise-la-Motte & Retheuil. — Type age: Eocene (Ypresian). — Current status: Diodora distans (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86) .

elegans , Fissurella – Deshayes, 1861-DescrCoqFoss: 240, 7, pl. 7, figs 17-20. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Current status: Diodora elegans (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 42).

elegans , Rimula – Deshayes, 1861-DescrCoqFoss: 245, 4, pl. 4, figs 1-4. — Type localities: Chaussy & Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32385, syntype from Chaussy. — Current status: Rimula elegans Deshayes, 1861 ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 42).

fenestrata , Emarginula – Deshayes, 1861-DescrCoqFoss: 250, 3, pl. 3, figs 37-41. — Type localities: Chaumont-en-Vexin & Lizy-sur-Ourcq. — Type age: Eocene (Lutetian-Bartonian). — Type material: UCBL-EM 30604, syntype. The specimen of Bartonian from Lizy-sur-Ourcq, UCBL-EM 30605 is not the same species. — Current status: Clypidina (Montfortula) fenestrata (Deshayes, 1861) ( Le Renard & Pacaud 1995: 87) . Clypidina (Montfortia) fenestrata (Deshayes, 1861) ( Pacaud 2008: 41) . Synonym of Montfortia elegans (Defrance, 1819) [ Emarginula ] (Pacaud herein).

fenestrella , Emarginula – Deshayes, 1863: 49-50, pl. 7, figs 12- 15, on pl. as Emarginula “ fenestrata ”; non E. fenestrella Dubois de Montpéreux, 1831 ( Emarginula fenestrata would be non Deshayes, 1861). — Type locality: La Réunion. — Type age: Recent. — Current status: Emarginula “ fenestrata ” Deshayes, 1863 ( Herbert 1987: 12, fig. 43; Herbert 2015: 21). Emarginula apolonia Pacaud, 2015b , replacement name for E. fenestrella Deshayes.

grata , Fissurella – Deshayes, 1861-DescrCoqFoss: 241, 7, pl. 7, figs 5-8. — Type locality: Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Type species (M) of Atractotrema Cossmann, 1888 . — Current status: Atractotrema gratum (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 41).

imbrex , Fissurella – Deshayes, 1861-DescrCoqFoss: 239, 20, pl. 20, figs 31-33. — Type localities: Mouchy-le-Châtel & Mouy. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32463, syntype from Mouchy-le-Châtel. — Current status: Diodora imbrex (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 42).

incerta , Fissurella – Deshayes, 1861-DescrCoqFoss: 237, 7, pl. 7, figs 25-27. — Type locality: Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32392, syntypes. — Current status: Diodora incerta (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 42).

intorta , Rimula – Deshayes, 1861-DescrCoqFoss: 245-246, 3, pl. 3, figs 29-32. — Type localities: Thiverval-Grignon & Chaussy. — Type age: Eocene (Lutetian). Rimula (Rimula) intorta Deshayes, 1861 ( Glibert 1962a: 11). — Current status: synonym of Rimularia dubia (Defrance, 1819) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 41).

magnifica , Fissurella – Deshayes, 1861-DescrCoqFoss: 236-237, pl. 8, figs 16-18. — Type locality: Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32395, syntypes. — Current status: Diodora magnifica (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 42).

mitis , Fissurella – Deshayes, 1830b-EncyMeth: 136. — Type locality: Faluns de Touraine & Italy. — Type age: Miocene. — Current status: synonym of Diodora multifida (Deshayes, 1830) ( Glibert 1949: 28; Landau et al. 2017: 82).

multifida , Fissurella – Deshayes, 1830b-EncyMeth: 136. — Type locality: Angers. — Type age: Miocene. — Current status: Diodora multifida (Deshayes, 1830) ( Landau et al. 2017: 82, pl. 4, fig. 1).

neglecta , Fissurella – Deshayes, 1830b-EncyMeth: 138; 1833b: 232; 1835b: 134; 1836-HistNatAnim2: 601; 1843-HistNatAnim3: 221; Guérin-Méneville & Deshayes 1868: [45], pl. 22, fig. 5, fig. of animal only. — Type locality: Mediterranean; Mr. Dusgate. — Type age: Recent. — Type material:MNHN-IM-2000-4789, 6 syntypes. — Current status: synonym of Diodora italica (Defrance, 1820) [ Fissurella ] ( Sabelli et al. 1990: 124).

ornata , Emarginula – Deshayes, 1825-DictClass: pl. [77], fig. 2; 1831-DictClass17: 118. — Type locality: none given. — Type age: none given. Preoccupies E. ornata Millet de La Turtaudière, 1865 , which was renamed Emarginula octaviana Coen, 1839 . — Current status: taxon inquirendum.

radiola, Fissurella – Deshayes, 1830b-EncyMeth: 136; 1836-HistNatAnim2: 600-601; 1843-HistNatAnim3: 221. — Type locality: Malouines [ Falkland Islands]; Lesson. — Type age: Recent. — Type material: MNHN-IM-2000-4796, syntype. — Current status: synonym of Fissurella picta (Gmelin, 1791) [ Patella ] (D. Zelaya, pers. comm., 2021).

rudis , Fissurella – Deshayes, 1830b-EncyMeth: 133, non Fissurella rudis (Röding, 1798) [ Patella ]; 1836-HistNatAnim2: 601-602; 1843-HistNatAnim3: 221. — Type locality: Paita, Peru; Lesson. — Type age: Recent.– Type material:MNHN-IM-2000-4801, lectotype ( McLean, 1984); MNHN-IM-2000-4802, 4 paralectotypes. — Current status: synonym of Fissurella costata Lesson, 1831 ( McLean 1984: 34, figs 95-108).

scutellata , Emarginula – Deshayes, 1863: 45-46, pl. 7, figs 1-2. — Type locality: La Réunion. — Type age: Recent. — Type material: MNHN-IM-2000-4806, lectotype; MNHN-IM-2000-4807; MNHN-IM-2000-4808, 3 paralectotypes. Hemitoma (Hemitoma) scutellata (Deshayes, 1863) ( Herbert 1987: 1-3, figs 1-5). — Current status: Octomarginula scutellata (Deshayes, 1863) ( McLean 2011: 415; Herbert 2015: 26).

spinosa , Emarginula – Deshayes, 1863: 48-49, pl. 7, figs 9-11. — Type locality: La Réunion. — Type age: Recent. Emarginula spinosa Deshayes, 1863 . — Current status: synonym of Emarginula decorata Deshayes, 1863 (Manousis & Galinou-Mitsoudi 2014: 3, fig. 2a).

squamosa , Fissurella – Deshayes, 1824-DescrCoqFoss: 21, 1, pl. 2, figs 1-3. 1824 -DictClass6: 527; 1830b-EncyMeth: 135; 1836-HistNatAnim2: 603; 1843-HistNatAnim3: 222; 1861-DescrCoqFoss: 237. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). — Current status: Diodora squamosa (Deshayes, 1824) ( Le Renard & Pacaud 1995: 86; Pacaud 2008: 42; Courville et al. 2012: 58, fig. 2: 8, 60, pl. 2, fig. 8).

sublamellosa , Fissurella –Deshayes, 1861-DescrCoqFoss: 238, 7, pl. 7, figs 21-24. — Type localities:Cuise-la-Motte, Aizy-Jouy, Laon & Hérouval. — Type age: Eocene (Ypresian).— Type material: UCBL-EM 32391, syntypes from Cuise-la-Motte. — Current status: Diodora sublamellosa (Deshayes, 1861) ( Le Renard & Pacaud 1995: 86) .

subrotunda , Fissurella –Deshayes, 1830b-EncyMeth: 135; 1836-HistNatAnim2: 602; 1843-HistNatAnim3: 221. — Type locality: Peru. — Type age: Recent. — Type material: MNHN-IM-2000-4809, 8 syntypes. — Current status: synonym of Fissurella peruviana Lamarck, 1822 ( McLean 1984: 21-23, figs 31-50) .

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macrochisma , Fissurella – Deshayes, 1830b-EncyMeth: 132-133, as a Sowerby species; 1836-HistNatAnim2: 603, ex Chemnitz ms. — Type locality: “ New Zealand ”. — Type age: Recent. 1843-HistNatAnim3: 222. Listed by Sherborn (1928: 3774) as an 1836 Deshayes species, this fissurellid had its origin as Patella macroschisma [Lightfoot], 1786, who cited a still-earlier figure. It is the type species (M) of Macroschisma J. E. Gray, 1835 . Although the type species itself is a nomen dubium , there are several well documented species in this genus.

scutulata, Emarginula – G. B. Sowerby II, 1873: pl. 5, fig. 37, ex Deshayes ms. It is unclear whether this species, without a locality, was a misspelling of Emarginula scutellata Deshayes, 1863 , from La Réunion (see above), or if it was a new species ( Petit 2009: 165, 188, note 9). — Current status: taxon inquirendum.