HELICIDAE Rafinesque, 1815

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 692-694

Family HELICIDAE Rafinesque, 1815

arvernensis , Helix – Deshayes, 1863-DescrCoqFoss: 812. Introduced for previously misidentified Helix cocquii Brongniart, 1810 , by Bouillet (1835) and Boissy (1844). — Type localities: Machal, Chaptuzat, Jussat, Puy de Marman, Nonette & Roche-Blanche. — Type age: Oligocene (Chattian). Noulet (1858: 420) had earlier introduced the new name Helix obesa for the species misidentified by Bouillet and de Boissy. However, this name is a junior primary homonym of Helix obesa Beck, 1837 . Synonym of Cepaea moroguesi (Brongniart, 1810) [ Helix ] ( Truc 1971: 281). — Current status: Palaeotachea moroguesi (Brongniart, 1810) ( Kadolsky 1989) .

asperula , Helix – Deshayes, 1832-EncyMeth: 251; 1838-HistNatAnim2: 137; 1843-HistNatAnim3: 317. — Type locality: Touraine. — Type age: Miocene. — Type species (OD) of Trachytachea Pfeffer, 1930 . Hemicycla asperula (Deshayes, 1832) (Wenz 1923: 573) . Helix (Eobania) turonensis [Deshayes] asperula Deshayes, 1830 ( Peyrot 1932: 433-434, pl. 16, figs 7-9). — Current status: synonym of Megalotachea turonensis (Deshayes, 1832) ( Truc 1971: 281) .

aureliana, Helix – Deshayes, 1863-DescrCoqFoss: 807-808, 51, pl. 51, figs 8-10, ex Brongniart ms. — Type localities: Marigny, Fay-aux-Loges, Neuville-aux-Bois, La Chapelle, Pontournois near Pithiviers & near Chartres. — Type age: Miocene (Aquitanian). — Current status: synonym of Cepaea moroguesi (Brongniart, 1810) (Wenz 1923: 639) , now Palaeotachea moroguesi (Brongniart, 1810) (MolluscaBase) .

barrandii , Helix – Deshayes, 1863-DescrCoqFoss: 808, 51, pl. 51, figs 19-21. — Type localities: Marigny, Fay-aux-Loges, La Chapelle & Chartres. — Type age: Oligocene (Rupelian and Chattian). — Current status: synonym of Cepaea moroguesi (Brongniart, 1810) [ Helix ] (Wenz 1923: 639), now Palaeotachea moroguesi (Brongniart, 1810) (MolluscaBase) .

beyrichi , Helix – Deshayes, 1863-DescrCoqFoss: 810-811, 52, pl. 52, figs 1-4. — Type locality: Fontainebleau. — Type age: Oligocene (Rupelian). — Current status: Cepaea beyrichi (Deshayes, 1863) (Wenz 1923: 608) .

brongniarti , Helix – Deshayes, 1863-DescrCoqFoss: 812-813, 51, as H. brongnarti , pl. 51, figs 14, 17-18 [fig. 14 not listed in text]. — Type localities: Pontournois, Fay-aux-Loges & Marigny. — Type age: Miocene (Aquitanian). 1865-DescrCoqFoss: 666 [correction noted]. — Current status: synonym of Cepaea moroguesi (Brongniart, 1810) (Wenz 1923: 639) , now Palaeotachea moroguesi (Brongniart, 1810) (MolluscaBase) .

cingulina , Helix – Deshayes, 1840-HistNatTerrFluv1: 26;1851-HistNatTerrFluv: 13, pl. 68, fig. 4. Uncaptioned pl. 68 issued in 1832. — Type locality: “Scheneberg” [Schneeberg], Austria. — Type age: Recent. Preoccupies Helix cingulina Strobel, 1844 . — Current status: Chilostoma achates cingulina (Deshayes, 1839) ( Schileyko 2013: 154) .

cyclolabris , Helix –Deshayes,1840-HistNatTerrFluv1: 32-33; 1851-HistNatTerrFluv: 13, pl. 68, fig. 7. Uncaptioned pl. 68 issued in 1832. — Type locality: Naxos Island , Greece. — Type age: Recent. — Current status: Thiessea cyclolabris (Deshayes, 1839) (MolluscaBase) .

defrancii , Helix – Deshayes, 1863-DescrCoqFoss: 804-805, 51, pl. 51, figs 11-13 [in text as figs 11-14]; 1863-DescrCoqFoss: 666 [correction noted]. — Type localities: Marigny, La Chapelle, Fay-aux-Loges, Neuville-aux-Bois, Blois & Montabuzard. — Type age: Miocene (Aquitanian). — Current status: synonym of Cepaea moroguesi (Brongniart, 1810) [ Helix ] (Wenz 1923: 639), now Palaeotachea moroguesi (Brongniart, 1810) (MolluscaBase) .

discordialis , Helix – Deshayes, 1839-HistNatTerrFluv1: 13-14; 1851-HistNatTerrFluv: 14, pl. 74, fig. 1, ex Férussac ms. Uncaptioned pl. 74 issued in 1832. — Type locality: not stated. — Type age: Recent. — Type material:MNHN-IM-2000-1863, holotype. — Type species (M) of Erigone Albers, 1850 , non Audoin, 1826 [ Arachnida]; Helerigone Strand, 1910 , replacement name, but now considered to be a synonym of Chloritis Beck, 1837 . — Current status: the Philippines Chloritis discordialis (Deshayes, 1839) (MolluscaBase) .

discus, Helix – Deshayes, 1850-HistNatTerrFluv1: 360-361; 1851-HistNatTerrFluv: 12, pl. 62A, fig. 4, non Thomä, 1845. — Type locality: none given. — Type age: Recent. — Type material: MNHN-IM-2000-2025, holotype. — Current status: synonym of Obba parmula (Broderip, 1841) [ Helix ] from the Philippine Islands (MolluscaBase).

dubia , Helix – Deshayes, 1824-DescrCoqFoss: 55, 3, pl. 6, fig. 3, non G. Fischer, 1807; 1832-EncyMeth: 250; 1863-DescrCoqFoss: 826 [species now doubtful]. — Type localities: Saint-Cyr & Isle of Wight. — Type age: Recent. — Current status: Taxon inquirendum (Wenz 1923: 1111) .

duvauxii, Helix – Deshayes,1832-EncyMeth: 251-252;1838-HistNatAnim2: 138; 1843-HistNatAnim3: 318. — Type locality:Touraine; Mr. Duvau. — Type age: Miocene. Synonym of Hemicycla asperula (Deshayes, 1832) (Wenz 1923: 574) . — Current status: synonym of Megalotachea turonensis (Deshayes, 1832) ( Truc 1971: 281) .

eumicron, Helix –Deshayes, 1863-DescrCoqFoss: 821-822, 52, pl. 52, figs 19-22. — Type locality: Côte Saint-Martin (Étampes). — Type age: Oligocene (Rupelian). — Type material: MNHN.F.J04143, syntype. — Current status: taxon inquirendum (Wenz 1923: 1111).

eversa, Helix – Deshayes, 1851-HistNatTerrFluv1: 395-396, 22, Foss pl. 1, figs 5-7. — Type locality: Faluns de Touraine. — Type age: Miocene. Synonym of – Current status: Megalotachea turonensis (Deshayes, 1831) [ Helix ] ( Höltke & Rasser 2016: 251).

exornata , Helix – Deshayes, 1850-HistNatTerrFluv1: 220-221, pl. 17A, figs 9-11; 1851-HistNatTerrFluv: 5, in fig. expl. as " H. ornata ". Uncaptioned pl. 17A issued in 1839. — Type locality: none given; Férussac coll. — Type age: Recent. — Current status: Leptaxis exornata (Pilsbry 1888-1889: 198, pl. 51, figs 13-14, as Helix (Leptaxis) .

filosa , Helix –Deshayes, 1850-HistNatTerrFluv1: 220;1851-HistNatTerrFluv: 10, pl. 29A, figs 22-23. — Type locality: none given. — Type age: Recent. — Current status: possibly Cepaea (Pilsbry 1895: 322, following L. Pfeiffer and as Helix ( Tachea W. Turton, 1831 ), a junior homonym, non J. Fleming, 1822.

gibbosula , Helix – Deshayes, 1850-HistNatTerrFluv1: 224; 1851- HistNatTerrFluv: 16, pl. 107, figs 1-3 [figs cited in the text in error as “1-2, 8”]. — Type locality: none given. — Type age: Recent. — Current status: possibly Cepaea (Pilsbry, 1895: 322, following L. Pfeiffer and as Helix ( Tachea W. Turton, 1831 ), subgenus a junior homonym, non J. Fleming, 1822. hongkongensis, Helix – Deshayes, 1874b: 3, pl. 3, figs 7-9; 1876: 87- 88. — Type locality: Hong Kong, China. — Type age: Recent. — Current status: taxon inquirendum.

insubrica, Helix –Deshayes, 1840-HistNatTerrFluv1: 30; 1851-HistNatTerrFluv: 13, pl. 68, fig. 3, non De Cristofori & Jan, 1832. Uncaptioned pl. 68 issued in 1832. The figure was not those cited in the text, pl. 69, figs 1-5, which were instead of Helix setipila Rossmässler, 1835 . — Type locality: Near Lake Como, Italy. — Type age: Recent. — Current status: synonym of the European Chilostoma tigrinum (De Cristofori & Jan, 1832) [ Helix ]. However, the latter is also a primary junior homonym, non Férussac, 1821, so a replacement name may be needed.

joncheryensis , Helix – Deshayes, 1858: 553, nomen nudum.

lucasii, Helix –Deshayes, 1850-HistNatTerrFluv1: 122-124;1851-HistNatTerrFluv: 16, pl. 96, figs 8-12. — Type locality: Oran, Algeria; sandy areas near sea. — Type age: Recent. — Type material:MNHN- IM-2000-31721, 10 syntypes;MNHN-IM-2000-31722, 3 syntypes; MNHN-IM-2000-31723, 2 syntypes;MNHN-IM-2000-31724, 3 syntypes. — Current status: synonym of Otala lactea (O. F. Müller, 1774) [ Helix ] ( Kneubühler et al. 2019: 17-19, fig. 16D).

maguntina , Helix – Deshayes,1832-EncyMeth: 252; 1838-HistNatAnim2: 138; 1843-HistNatAnim3: 318. — Type locality: Mayence, Germany. — Type age: Miocene; Mr. Hoeninghaus. — Type species (OD) of Mesotachea Pfeffer, 1930 . Megalotachea maguntina (Deshayes, 1832) (H. Nordsieck 2017: 68, fig. 13). — Current status: Palaeotachea maguntina (Deshayes, 1832) ( Kadolsky 2020: 16) .

munieri , Helix – Deshayes, 1863-DescrCoqFoss: 809, 51, pl. 51, figs 22-24. — Type locality: Côte Saint-Martin (Étampes). — Type age: Oligocene (Rupelian). — Type material: MNHN.F.J03356, holotype. Does not preoccupy Helix munieriana Crosse & Debeaux, 1863 , which was unnecessarily renamed Helix tchefouensis Crosse & Debeaux, 1864 ) ( Breure et al. 2022a: 165, 215). — Current status: Parachloraea munieri (Deshayes, 1863) (Lozouet et al. 2012: 416- 417, fig. 282: 5-8).

noueli , Helix – Deshayes, 1863-DescrCoqFoss: 803-804, 51, pl. 51, figs 15-16. — Type localities: Montabuzard, La Chapelle, near Chartres & Della Badia (Piémont). — Type age: Eocene (Lutetian). — Type material: MNHN.F.J04129, syntype. — Current status: Cepaea noueli (Deshayes, 1863) (Wenz 1923: 647) .

obscura , Helix – Deshayes, 1874b: 13, pl. 3, figs 11-12; 1876: 86-87, non O. F. Müller, 1774. — Type locality: Hong Kong, China. — Type age: Recent. — Current status: taxon inquirendum .

ornata , Helix – see: exornata , Helix .

perforata , Helix – Deshayes, 1874b: 14, pl. 3, figs 29-32; 1876: 92- 93. — Type locality: Peking, China. — Type age: Recent. Preoccupies Helix niciensis var. perforata Caziot, 1910 . — Current status: taxon inquiredum .

plicatilis , Helix –Deshayes, 1870b: 20-21; 1874b: 13, pl. 2, figs 13- 15. — Type locality: Moupin, Tibet. — Type age: Recent. — Type material: MNHN-IM-2000-34195, 8 syntypes. — Current status: taxon inquirendum .

pouzolzi , Helix – Deshayes, 1831c: [1-2], pl. 30, ex Payraudeau ms; 1850-HistNatTerrFluv1: 59-61;1851-HistNatTerrFluv: 13, pl. 69G, figs 1-8 [in text as “1-6”]. — Type locality: Corsica. — Type age: Recent. — Type material:MNHN-IM-2000-31636, syntype. This species was a nomen nudum inPayraudeau (1826: 102-103). — Current status: Chilostoma (Dinarica) pouzolzi (Deshayes, 1830) ( Schileyko 2006: 1780-1781, fig. 2280, with this species mistakenly listed as the type species of Dinarica Kobelt, 1902; Schileyko 2013: 158). rupelli, Helix – Deshayes, 1870b: 19-20 [as rupelli]; 1874b: 12 [as ruppelli], pl. 2, figs 1-2. — Type locality: Moupin, Tibet [west of Chengdu, Sichuan, China]. — Type age: Recent. — Type material: MNHN-IM-2000-34193, syntype. Perhaps intended to honor Wilhelm Peter Eduard Simon Rüppell (1794-1884), a German biologist who explored northern Africa, but Deshayes did not say so, and his original spelling has to stand. — Current status: taxon inquirendum.

scabriuscula , Helix – Deshayes, 1832-EncyMeth: 258; 1838-HistNatAnim2: 135;1843-HistNatAnim3: 316. — Type locality: Sicily; Mr. Caron . — Type age: Recent. Murella scabriuscula (Deshayes, 1832) (K. L. Pfeiffer, 1931: 108-111, pl. 8, fig. 10). — Current status: Marmorana (Murella) scabriuscula (Deshayes, 1832) ( Schileyko 2013: 158) .

securiformis, Helix – Deshayes, 1850-HistNatTerrFluv1: 362; 1851-HistNatTerrFluv: 13, pl. 64A, figs 5-6. — Type locality: none given. — Type age: Recent. — Current status: taxon inquirendum.

sicana, Helix –Deshayes, 1838-HistNatAnim2: 130-131;1850-HistNatTerrFluv1: 244-245;1851-HistNatTerrFluv: 9, pl. 28B, fig. 7, ex Férussac ms. — Type locality: Sicily. — Type age: Recent. — Current status: Marmorana (Murella) platychela [ Menke, 1830] sicana (Deshayes, 1838) (K. L. Pfeiffer 1931: 106-107, pl. 9, fig. 32, as “Fer[ussac].”).

subechinata, Helix – Deshayes, 1870b: 22; 1874b: 13, pl. 2, figs 19- 21. — Type locality: Moupin, Tibet. — Type age: Recent. — Type material: MNHN-IM-2000-34198, 2 syntypes. — Current status: taxon inquirendum .

sudetica, Helix – Deshayes, 1850-HistNatTerrFluv1: 41; 1851-HistNatTerrFluv: 14, pl. 69J, figs 18-19, ex Charpentier ms. — Type localities: Silésie, Warth & near Neiffe; Mr. Audouin. Uncaptioned pl. 69J issued in 1841. — Type age: Recent. — Current status: synonym of Faustina faustina ( Rossmässler, 1835) [ Helix ] (R. Bank, pers. comm., November 23, 2022), type species of Faustina Kobelt, 1904 , by tautonomy.

tessellata , Helix – Deshayes, 1850-HistNatTerrFluv1: 232-233; 1851-HistNatTerrFluv: 5, pl. 17A, figs 12-13, ex Férussac ms. Uncaptioned pl. 17A issued in 1839. — Type localities: Majorca & Baléres. — Type age: Recent. — Current status: synonym of Allognathus grateloupi (Graells, 1846) [ Helix ] (R. Bank, pers. comm., November 23, 2022).

turonensis , Helix – Deshayes, 1831b: 139-140, 259, pl. 1, figs 1-2; 1832-EncyMeth: 251.1838-HistNatAnim2: 137-138;1843-HistNatAnim3: 317-318;1851-HistNatTerrFluv1: 394-395, 22, Foss. pl. 1, figs 2, 4. — Type locality: Mayence. — Type age: Miocene. — Type species (SD Truc, 1971) of Megalotachea Pfeffer, 1930 . — Current status: Megalotachea turonensis (Deshayes, 1831) ( Harzhauser et al. 2014b: 37, pl. 11, figs 1-4).

volhyniensis, Helix – Deshayes, 1840-HistNatTerrFluv1: 35, ex Andrzejowski ms. In synonymy with Helix faustina Rossmässler, 1835 , ex Ziegler ms. — Current status: synonym of the European Faustina faustina ( Rossmässler, 1835) .

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chastellii , Helix – This species was credited to “Deshayes in Férussac, 1832”, in Fischer-Piette et al. (1994: 230), but Deshayes was not involved when Férussac made this name available, although Deshayes did discuss it in his part of this work in 1850. — Current status: Kalidos chastellii (Férussac, 1832) .

chemnitziana , Clausilia – Deshayes, 1838-HistNatAnim2: 217, ex Férussac ms. Jamaica. — Type age: Recent. Listed by Sherborn (1925: 1221) as a Deshayes name. However, as Deshayes made clear, this name was proposed by Férussac (1821), as Helix (Cochlodina) chemnitziana , also listed by Sherborn. — Current status: Apoma chemnitzianum ( Férussac, 1821) .

Unallocated “ Helix ”

atrolineata, Helix – Deshayes, 1851-HistNatTerrFluv: 10, pl. 29A, figs 4-7; not in text. — Type locality: none given. — Type age: Recent. — Current status: taxon inquirendum.

complanata , Helix – Deshayes, 1839-HistNatTerrFluv1: 9-10; 1851-HistNatTerrFluv: 13, pl. 69B, fig. 2, non Linnaeus, 1758. — Type locality: Bolivia. — Type age: Recent. — Current status: taxon inquirendum .

nicobarica , Helix – Deshayes, 1839: [1], pl. 3, ex Chemnitz ms. — Type locality: none given, but Nicobar Islands implied. — Type age: Recent. — Current status: taxon inquirendum.

perdita, Helix – Deshayes, 1876: 93-95, 99, pl. 1, figs 12-15, non Reeve, 1851. — Type locality: Peking, China. — Type age: Recent. — Current status: taxon inquirendum.

scalprum, Helix – Deshayes, 1850-HistNatTerrFluv1: 362-363; 1851-HistNatTerrFluv: 13, pl. 64A, figs 2-4 [in 1850 text in error as “pl. 62A”], ex Valenciennes ms. — Type locality: none given. — Type age: Recent. — Current status: taxon inquirendum.

semiparita, Helix – Deshayes, 1850-HistNatTerrFluv1: 187-188; 1851-HistNatTerrFluv: 15, pl. 89, fig. 5. — Type locality: none given. — Type age: Recent. — Current status: taxon inquirendum.

striatissima , Helix – Deshayes, 1876: 84, 99, pl. 1, figs 1-4. — Type locality: Hong Kong. — Type age: Recent. — Current status: taxon inquirendum.

villae , Helix –Deshayes, 1850-HistNatTerrFluv1: 295-296, pl. “91” [90], fig. 9; 1851-HistNatTerrFluv: 15. — Type locality: none given. — Type age: Recent. Preoccupies Helix villae L. Pfeiffer, 1856 , which has been synonymized with Oxychilus mortilleti (L. Pfeiffer, 1859) [ Helix ]. — Current status: taxon inquirendum.