ACAVIDAE Pilsbry, 1895

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 686

Family ACAVIDAE Pilsbry, 1895

atropos , Helix –Deshayes, 1850-HistNatTerrFluv1: 56-57, ex Férussac ms; 1851-HistNatTerrFluv: 14, pl. 69H, figs 13-14. Uncaptioned pl. 69H issued in 1840. — Type locality: near Diego-Suarez Bay, Madagascar. — Type age: Recent. — Current status: Ampelita atropos (Deshayes, 1850) ( Fischer-Piette et al. 1994: 102-103, pl. 8, figs 14- 16; Emberton & Rakotondrazafy 2020: 177-179, 182, figs 80-87).

clotho , Helix –Deshayes, 1850-HistNatTerrFluv1: 57-58, ex Férussac ms; 1851-HistNatTerrFluv: 14, pl. 69I, fig. 3. Uncaptioned pl. 69I issued in 1840. — Type locality: near Diego-Suarez Bay, Madagascar. — Type age: Recent. — Current status: Ampelita clotho (Deshayes, 1850) ( Fischer-Piette et al. 1994: 110-111, pl. 10, figs 9-11; Emberton & Rakotondrazafy 2020: 181-183, 178, figs 50-53).

consanguinea , Helix – Deshayes, 1850-HistNatTerrFluv1: 59, ex Férussac ms; 1851-HistNatTerrFluv: 13, pl. 69H, figs 1-2. Uncaptioned pl. 69H issued in 1840. — Type locality: near Diego Suarez Bay, Madagascar. — Type age: Recent. — Current status: Ampelita consanguinea (Deshayes, 1850) ( Fischer-Piette et al. 1994: 94-95, pl. 11, figs 13-15; Emberton & Rakotondrazafy 2020: 183, 190, figs 136-139).

echinophora , Helix – Deshayes, 1850-HistNatTerrFluv1: 287-288, ex Férussac ms; 1851-HistNatTerrFluv: pl. 10A, figs 7-9, ex Férussac ms. Uncaptioned pl. 10A issued in 1839. — Type locality: Madagascar. — Type age: Recent. This species name first appeared as a nomen nudum in L. Pfeiffer (1848: 434). — Type material: MNHN-IM-2000-2082, lectotype ( Groh & Poppe 2002). Helicophanta echinophora (Deshayes, 1850) ( Fischer-Piette et al. 1994: 163-164, pl. 25, figs 6-8, pl. 30, figs 6-7, text-figs 68-69, as “Férussac, 1839”. — Current status: Embertoniphanta echinophora (Deshayes, 1850) ( Groh & Poppe 2002: 43-45: pl. 27, figs 1-3, text-figs 16-17; Emberton & Rakotondrazfy 2020: 203-204, figs 77-79).

granulosa , Helix –Deshayes, 1850-HistNatTerrFluv1: 61; 1851-HistNatTerrFluv: 13, ex Férussac ms, pl. 69H, figs 7-10. Uncaptioned pl. 69H issued in 1840. — Type locality: near Diego-Suarez Bay, Madagascar; Goudot. — Type age: Recent. — Current status: Ampelita granulosa (Deshayes, 1850) ( Fischer-Piette et al. 1994: 114-115, pl. 12, figs 1-3; Emberton & Rakotondrazafy 2020: 187, 186, figs 107-110).

lachesis, Helix – Deshayes, 1850-HistNatTerrFluv1: 57; 1851-HistNatTerrFluv: 13, pl. 69H, figs 3-4, ex Férussac ms. Uncaptioned pl. 69H issued in 1840. — Type locality: near Diego-Suarez Bay, Madagascar; Goudot. — Type age: Recent. — Current status: Ampelita lachensis (Deshayes, 1850) ( Fischer-Piette et al. 1994: 108-109, pl. 10, figs 4-5; Emberton & Rakotondrazafy 2020: 191, 178, figs 46-49).

microdonta , Helix – Deshayes, 1832-EncyMeth: 266; 1839-HistNatTerrFluv1: 6; 1850: 214-215, as a synonym of H. unidentata ( Holten, 1802) ; 1851-HistNatTerrFluv: 14, pl. 72, fig. 13. — Type locality: Senegal. — Type age: Recent. — Current status: synonym of Stylodonta unidentata “Chemnitz ” [ Holten, 1802] [ Helix ] from the Seychelles ( Groh & Poppe 2002: 25; Bank & Menkhorst 2008: 970). Bieler & Slapcinsky (2000: 65) maintained that Deshayes’ treatment of 1839 was not the same as his original 1832 species.

viridis , Helix – Deshayes,1832-EncyMeth: 264;1838-HistNatAnim2: 102-103; 1843-HistNatAnim3: 305. — Type locality: Madagascar.— Type age: Recent. — Type material: MNHN-IM-2000-28794, 18 syntypes. — Current status: Eurystyla viridis (Deshayes, 1832) ( Fischer-Piette et al. 1994: 148-149, pl. 21, fig. 14, as “1838”; Groh & Poppe 2002: 56-57, pl. 41, figs 1-9, text-fig. 19, as “1838”).

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elongata , Megaspira – Deshayes, 1863-DescrCoqFoss: 864, 55, pl. 55, figs 13-15. Made available by Melleville (1843: 92-93 [46-47, 84], pl. 4, fig. 23-25), as Pupa elongata . — Type locality: Châlons-sur-Vesle. — Type age: Paleocene (Thanetian). Primary junior homonym of Pupa elongata Bouillet, 1836 . Kadolsky (2021) proposed to maintain the junior primary homonym under ICZN Code Article 23.9.3.

REMARK

Two species from Madagascar have been incorrectly attributed to Férussac in 1839 or 1840 in the plates of the continuation of his work that were actually issued by Deshayes. However, these plates were uncaptioned, with the accompanying text and plate explanations only appearing in 1850 and 1851. The first species had already been made available as Helix goudotiana L. Pfeiffer, 1845 ( Pfeiffer 1845: 155), also discussed in L. Pfeiffer (1847: 18), evidently copied from labels with Férussac material or ms notes. It is now considered a synonym of Embertoniphanta oviformis (Grateloup, 1840) [ Helix ] ( Emberton & Rakotondrazafy 2020: 202-206, figs 8-21).

A second species, Helix amphibulima , also from Madagascar, had appeared as a nomen nudum in Beck (1838: 46, ex Valenciennes ms) and in then L. Pfeiffer (1847: 18), as a synonym of H. oviformis Grateloup, 1840 ). However, used as an available name before 1961 by Crosse & P. Fischer, in Grandidier (1889) (ICZN Code Art. 11.6.1). This is thus Embertoniphanta amphibulima (L. Pfeiffer, 1847) ; it is the type species of the genus Embertoniphanta Groh & Poppe, 2002 .