Pseudosmittia navama, Ferrington Jr & Saether, 2011

Ferrington Jr, Leonard C. & Saether, Ole A., 2011, A revision of the genera Pseudosmittia Edwards, 1932, Allocladius Kieffer, 1913, and Hydrosmittia gen. n. (Diptera: Chironomidae, Orthocladiinae) 2849, Zootaxa 2849 (1), pp. 1-314 : 281-282

Pseudosmittia navama sp. n.

( Fig. 108 A–E )

Type material: Holotype male, USA: California, Contra Costa County, Richmond , salt marsh, 2.ii.1969, G. Grodhaus ( USNM) . – Paratypes, USA: California: 34 males, as holotype; Contra Costa County, 2.5 mi W. Pinole, salt marsh, 2 males, 7.iii. & 18.iv.1969, R. E. Doty; Monterey County, mouth of Carmel River , 8 males, as holotype., 21.iii.1967, G. Grodhaus; Alameda County , Oakland, golf course near airport, 7 males, 21.v.1969, B. Keh & R. E. Doty; Solano County , Vallejo, Lake Marie (?), 4 males, 12.v.1969, E. G. Mezger & G. Grodhaus; Alameda County , 1 male, 20.ix.1968, R. S. Lave; Sonora County , Bodega Bay , 1 male, 19.x.1947, W. W. Wirth; Marin County , San Quentin Point, 1 male, 31.iii.1949, L. W. Quate ( CAL, CNC, JES, UMSP, USNM, ZMBN) . Oregon: Clatsop Co., Ft. Stevens State Park, parking lot of South Jetty overlook, 6 males, 23.x.1994, M. Spies ( ZMBN) .

Diagnostic characters: P. navama differs from the other members of the subgroup except P. parinavama by having a well developed wing anal lobe, a sinuate Cu 1, and conspicuous microtrichia to apex of a parallel-sided anal point. From P. parinavama it differs by having no setae on R and R 4+5, at most 1 seta on R 1, and only 3–5 sensilla clavata at apex of the third palpomere.

Etymology: From the Latin navis, a ship, boat, and hamas, a hook, referring to the boat-hook- like shape of the inferior volsella.

Male imago (n = 10, except when otherwise stated)

Total length 2.14–2.84, 2.50 mm. Wing length 1.17–1.63, 1.45 mm (63). Total length/wing length 1.55–1.93, 1.75. Wing length/length of profemur 2.73–2.97, 2.87. Coloration yellowish brown with slightly darker vittae.

Head ( Fig. 108 B ). AR 0.73–0.97, 0.78 (53). Terminal flagellomere 270–360, 320 µm (55) long, with 30–46, 36 µm long strong apical seta. Temporal setae 6–10, 7, consisting of 2–6, 3 inner verticals, 3–5, 4 outer verticals and 0–1, 0 postorbitals. Clypeus with 6–9, 7 setae. Tentorium 128–158, 143 µm long, 30–37, 32 µm wide. Stipes 110–146, 125 µm long, 18–37, 26 µm wide. Palpomere lengths (in µm): 25–32, 29; 32–41, 38; 69–91, 81; 66–98, 82; 96–133, 117. Third palpomere ( Fig. 108 C ) with 3–5, 4 slightly scalpellate sensilla clavata.

Thorax. Antepronotal lobes only slightly reduced medially, antepronotum with 1–3, 1 lateral setae. Dorsocentrals 11–15, 13; acrostichals 2; prealars 4–5, 4. Scutellum with 6–11, 8 setae.

Wing ( Fig. 108 A ). VR 1.40–1.48, 1.44. Anal lobe projecting. Costal extension 39–69, 53 µm long. R 4+5 ending slightly proximal to apex of M 3+4; Cu 1 sinuate. Brachiolum with 1 (65) seta, R with 0–1, 0 (65) setae, other veins bare.

Legs. Spur of front tibia 39–48, 45 µm long; spurs of middle tibia 23–32, 28 µm and 14–23, 19 µm long; of hind tibia 46–53, 48 µm and 18–23, 20 µm long. Width at apex of front tibia 25–32, 29 µm; of middle tibia 27–34, 30 µm; of hind tibia 41–50, 45 µm. Comb of 12–16, 14 setae; shortest setae 25–32, 28 µm long; longest setae 39–55, 46 µm long. Lengths and proportions of legs as in Table 76.

Hypopygium ( Fig. 108 E ). Anal point 21–46, 32 µm (63) long, with conspicuous microtrichia to apex; tergite IX with 7–16, 11 setae; laterosternite IX with 4–5, 5 setae. Phallapodeme 71–91, 80 µm long; transverse sternapodeme 46–57, 52 µm long, no oral projections. Virga ( Fig. 108 D ) 15–27, 20 µm long. Gonocoxite 158–194, 172 µm long; superior volsella single, digitiform, with rounded apex, 18–32, 27 µm (63) long, 13–25, 18 µm (63) wide; inferior volsella boathook-shaped, 18–34, 24 µm (63) long, 13–23, 17 µm (63) wide. Gonostylus 69–82, 75 µm long; megaseta 14–17, 16 µm long. HR 2.12–2.58, 2.31; HV 3.10–3.56, 3.35.

Female, pupa and larva

Unknown.

Distribution

The species is only known from eight localities in California and one in Oregon, USA .

Figures

FIGURE 108. Pseudosmittia spp., males. A–E. P. navama sp. n. A—wing; B—head; C—third palpomere; D—virga; E— hypopygium, left dorsal aspect, right ventral aspect. F—J. P. parinavama sp. n. F—wing; G—head; H—third palpomere; I— virga; J—hypopygium, left dorsal aspect, right ventral aspect.