Hoffmannia dwyeri W.C.Burger

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Hoffmannia dwyeri W.C.Burger Hoffmannia nicotianifolia (M.Martens & Galeotti)

Terrestrial herbs or shrubs. Distribution in PILA 1300– L.O.Williams

2100 m, mixed forest in transition to cloud forest, cloud Terrestrial herbs to shrubs. Distribution in PILA 1100–

forest, low elevation oak forest. Global distribution: 2200 m, mixed forest in transition to low elevation forest,

Endemic to Panama. Records: B. Hammel 5814 (MO). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Mexico

Hoffmannia hondurensis Standl. to Colombia. Records: C. Hamilton 3899 (MO), C.

Terrestrial herbs to shrubs. Distribution in PILA 1100– Hamilton 771 (MO), J. A. Duke 13634 (MO), K. E. Blum

1100 m, mixed forest in transition to low elevation forest. 2619 (MO), L.D. Gómez 23063 (MO).

Global distribution: Honduras, Panama. Records: D.

Santamaría 7901 (INB, PMA). Hoffmannia pittieri Standl. Terrestrial shrubs. Distribution in PILA 700–2400 m, low

Hoffmannia liesneriana L.O.Williams elevation mixed forest, mixed forest in transition to low

Terrestrial herbs or shrubs. Distribution in PILA 0–1100 elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution:

Costa Rica to Colombia. Records: E. Mora 305 (INB), G. m, low elevation mixed forest, mixed forest in transition Herrera 5981 (CR, INB, MO), J. A. Duke 13651 (MO), to low elevation forest, mixed forest in transition to cloud J. Dwyer 525 (MO), J. Dwyer 8765 (MO), J. Dwyer 8783 forest. Global distribution: Nicaragua to Peru. Records: (MO), J. Dwyer 8790 (MO), J. Dwyer 8797 (MO), J. A.K. Monro 4750 (BM, INB, MO), A. De Sedas 677 (INB, H. Kirkbride Jr. 147 (MO), K. E. Blum 2647 (MO), M. PMA), D. Santamaría 7439 (INB, PMA). Chinchilla 176 (INB), T. Croat 27005 (MO), T. Cochrane 6244 (MO). Ladenbergia brenesii Standl. Agujilla, Ascá, Quina, Tabaquillo de montaña, Zapatero

Hoffmannia psychotriifolia (Benth.) Griseb. (CR) . Terrestrial herbs or shrubs. Distribution in PILA 300–2400 Terrestrial trees. Distribution in PILA 900–2500 m, m, low elevation mixed forest, mixed forest in transition mixed forest in transition to low elevation forest, mixed to low elevation forest, mixed forest in transition to cloud forest in transition to cloud forest, cloud forest, low forest, cloud forest, low elevation oak forest. Global elevation oak forest, pasture. Global distribution: Costa distribution: Mexico to Panama, Greater Antilles, Lesser Rica to Panama. Records: A.K. Monro 6202 (PMA), N. Antilles. Records: A. Rodríguez 10611 (BM, INB, MO, Zamora 4054 (BM, INB, MO, PMA). PMA), A. Rodríguez 11937 (BM, INB, MO, PMA), C. Hamilton 3645 (MO), C. Hamilton 3901 (MO), D. Manettia barbata Oerst. Santamaría 7254 (BM, INB, MO, PMA), D. Solano 3896 Terrestrial twiners. Distribution in PILA 600–3400 m, (BM, INB, MO, PMA), D. Solano 5063 (BM, INB, MO, low elevation mixed forest, mixed forest in transition to PMA), D. Solano 5166 (BM, INB, MO, PMA), D. Solano low elevation forest, mixed forest in transition to cloud 5263 (BM, INB, MO, PMA), G. Davidse 23761 (CR, forest, cloud forest, low elevation oak forest, páramo. MO), G. Davidse 28414 (CR, MO), J. Dwyer 518 (MO), Global distribution: Costa Rica to Colombia. Records: M. K. E. Blum 2626 (MO), M. Alfaro 33 (INB), N. Zamora Grayum 10305 (INB, MO). 4064 (BM, INB, MO, PMA), N. Zamora 4140 (BM, INB, MO, PMA), N. Zamora 4178 (BM, INB, MO, PMA). Manettia flexilis Brandegee Terrestrial twiners. Distribution in PILA 1300–1800

Hoffmannia subauriculata Standl. m, mixed forest in transition to cloud forest. Global Terrestrial herbs to shrubs. Distribution in PILA 100–1700 distribution: Mexico to Ecuador. Records: A. Rodríguez m, low elevation mixed forest, mixed forest in transition 11949 (BM, INB, MO, PMA). to low elevation forest, mixed forest in transition to cloud forest. Global distribution: Costa Rica to Peru. Records: Nertera granadensis (Mutis ex L.f.) Druce A. Rodríguez 10731 (BM, INB, MO, PMA), D. Solano Frutilla del diablo (CR). 4331 (BM, INB, MO, PMA), D. Solano 4374 (BM, INB, Terrestrial herbs. Distribution in PILA 200–3500 m, MO, PMA), D. Solano 4460 (BM, INB, MO, PMA), G. low elevation mixed forest, mixed forest in transition to Davidse 23788 (CR, MO), G. Herrera 3124 (MO), G. low elevation forest, mixed forest in transition to cloud Herrera 5477 (INB, MO), L.D. Gómez 23148 (MO), V. forest, cloud forest, low elevation oak forest, páramo. Dryer 1092 (MO). Global distribution: Mexico to Argentina, Chile, Greater Antilles. Records: A. Fernández 708 (INB), A. Fernández

Hoffmannia subcapitata C.M.Taylor 958 (CR, INB, MO), A.K. Monro 4964 (BM, INB, MO), Terrestrial herbs. Distribution in PILA 300–1400 m, low A.K. Monro 5174 (BM), A.K. Monro 5277 (BM, INB, elevation mixed forest, mixed forest in transition to low MO), A.K. Monro 5603 (BM, INB, MO, PMA), A. elevation forest, mixed forest in transition to cloud forest. Rodríguez 11010 (BM, INB, MO, PMA), A. Soto 296 Global distribution: Costa Rica to Peru. Records: A. (INB), B.B. Klitgaard 785 (BM, INB, MO), D. Solano Rodríguez 11457 (INB), D. Santamaría 6567 (BM, INB), 3873 (BM, INB, MO, PMA), E. Alfaro 4681 (INB), G. D. Solano 4898 (BM, INB). Davidse 23980 (CR, MO), G. Davidse 25133 (CR, MO), G. Davidse 25209 (CR, MO), G. Davidse 26011 (CR,

Hoffmannia valerioi Standl. MO ), G. Davidse 26161 (CR, MO), G. Davidse 28691 Terrestrial herbs to shrubs. Distribution in PILA 100–2500 (CR, MO), G. Davidse 28969 (CR, MO), F. Quesada m, low elevation mixed forest, mixed forest in transition 1494 (CR, INB), L. Angulo 190 (INB), M.A. Thomas to low elevation forest, mixed forest in transition to cloud 1577 (MO). forest, cloud forest, low elevation oak forest. Global distribution: Costa Rica, Panama. Records: A. De Sedas Notopleura aggregata (Standl.) C.M.Taylor 941 (INB, PMA). Terrestrial herbs to shrubs. Distribution in PILA 1100– 2600 m, mixed forest in transition to low elevation forest,

Hoffmannia viridis Rusby mixed forest in transition to cloud forest, cloud forest, Terrestrial herbs to shrubs. Distribution in PILA 100–1500 low elevation oak forest, high elevation oak forest, dense thicket. Global distribution: Costa Rica to Colombia. Monro 4795 (BM, MO). Records: A. Fernández 645 (CR, INB, MO), A. Fernández 899 (CR, INB, MO), D. Santamaría 6690 (BM, INB, Notopleura elegans C.M.Taylor MO, PMA ), D. Solano 5159 (BM, INB, MO, PMA), G. Terrestrial herbs or terrestrial or facultatively epiphytic Davidse 29075 (MO), G. Herrera 5470 (INB, MO), G. shrubs. Distribution in PILA 1400–2500 m, mixed forest Herrera 5490 (INB, MO), M. Grayum 10291 (INB, MO), in transition to cloud forest, cloud forest, low elevation oak R. Villalobos 253 (INB). forest. Global distribution: Costa Rica, Panama. Records: A. Fernández 711 (CR, INB, MO), A. Fernández 976 (CR,

Notopleura amicitiae C.M.Taylor INB, MO ), A.K. Monro 5488 (BM, INB, MO, PMA), A. Terrestrial herbs or shrubs. Distribution in PILA 1300– Rodríguez 10845 (BM, INB, MO, PMA), D. Solano 3936 2200 m, mixed forest in transition to cloud forest, cloud (BM, INB, MO, PMA), J. Bittner 1801 (INB, MO). forest, low elevation oak forest. Global distribution: Costa Rica to Colombia. Records: B. Gamboa 630 (INB), Notopleura guadalupensis (DC.) C.M.Taylor D. Solano 5059 (BM, INB, MO, PMA), D. Solano 5150 Epiphytic shrubs. Distribution in PILA 1200– 1200 m, (BM, INB, MO, PMA), E. Alfaro 2267 (INB), G. Davidse mixed forest in transition to low elevation forest. Global 28445 (MO), L. Angulo 321 (INB). distribution: Mexico to the Guyanas, Greater Antilles. Records: G. Herrera 5380 (INB, MO).

Notopleura anomothyrsa (K.Schum. & Donn.Sm.) C.M.Taylor Notopleura longipedunculoides (C.M.Taylor) C.M.Taylor Terrestrial shrubs. Distribution in PILA 0–1000 m, low Terrestrial shrubs. Distribution in PILA 700– 700 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest. Global distribution: Costa Rica to elevation forest. Global distribution: Mexico to Ecuador. Ecuador. Records: G. Herrera 3255 (INB, MO). Records: A. Rodríguez 11616 (BM, INB, MO, PMA), D. Solano 4895 (BM, INB, MO, PMA), G. Herrera 5315 Notopleura maxonii (Standl.) C.M.Taylor (MO) . Epiphytic shrubs. Distribution in PILA 1200–2200 m, mixed forest in transition to low elevation forest, mixed

Notopleura camponutans (Dwyer & Hayden) C.M.Taylor forest in transition to cloud forest, cloud forest, low Terrestrial herbs or shrubs. Distribution in PILA 300–1700 elevation oak forest, pasture. Global distribution: Costa m, low elevation mixed forest, mixed forest in transition Rica, Panama. Records: A. Rodríguez 10674 (BM, INB, to low elevation forest, mixed forest in transition to cloud MO, PMA), A. Rodríguez 11530 (BM, INB, MO, PMA), forest. Global distribution: Costa Rica to Colombia. D. Santamaría 6652 (BM, INB, MO, PMA). Records: A. De Sedas 689 (PMA). Notopleura pacorana C.M.Taylor

Notopleura capacifolia (Dwyer) C.M.Taylor Terrestrial shrubs. Distribution in PILA 1000– 1000 m, Terrestrial herbs. Distribution in PILA 0–2100 m, low mixed forest in transition to low elevation forest,, dense elevation mixed forest, mixed forest in transition to low thicket. Global distribution: Endemic to Panama. Records: elevation forest, mixed forest in transition to cloud forest, A.K. Monro 4767 (BM). cloud forest, low elevation oak forest. Global distribution: Nicaragua to Peru. Records: A. Rodríguez 11272 (BM, Notopleura panamensis (Dwyer) C.M.Taylor INB, MO, PMA ), D. Solano 4360 (BM, INB, MO, PMA), Terrestrial shrubs or trees. Distribution in PILA 800–2200 S. Bridgewater 4144 (BM, INB, MO, PMA). m, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest,

Notopleura capitata C.M.Taylor low elevation oak forest. Global distribution: Costa Rica, Terrestrial herbs or shrubs. Distribution in PILA 0–1700 Panama. Records: J. Dwyer 7711 (MO). m, low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud Notopleura peperomiae (Standl.) C.M.Taylor forest, cloud forest. Global distribution: Nicaragua to Epiphytic shrubs. Distribution in PILA 600–1800 m, low Colombia. Records: D. Solano 3824 (BM, INB, MO, elevation mixed forest, mixed forest in transition to low PMA). elevation forest, mixed forest in transition to cloud forest. Global distribution: Costa Rica, Panama. Records: G.

Notopleura costaricensis C.M.Taylor Herrera 2688 (INB, MO). Terrestrial herbs or shrubs. Distribution in PILA 1000– 2300 m, mixed forest in transition to low elevation forest, Notopleura pithecobia (Standl.) C.M.Taylor mixed forest in transition to cloud forest, cloud forest, low Epiphytic shrubs. Distribution in PILA 600–1900 m, low elevation oak forest. Global distribution: Costa Rica to elevation mixed forest, mixed forest in transition to low Colombia. Records: A.K. Monro 4222 (BM, MO), A.K. elevation forest, mixed forest in transition to cloud forest. Global distribution: Costa Rica to Peru. Records: A. low elevation oak forest, páramo. Global distribution: Fernández 787 (CR, INB, MO), A.K. Monro 4246 (BM, Nicaragua to Panama. Records: E. Alfaro 3981 (INB), G. INB, MO), A.K. Monro 5522 (BM, INB, MO, PMA), A.K. Davidse 23872 (CR, MO), G. Davidse 28596 (CR, MO), Monro 5608 (BM, INB, MO, PMA), A.K. Monro 6209 G. Herrera 5450 (CR, INB, MO), M.A. Thomas 1614 (PMA), A. Rodríguez 10580 (BM, INB, MO, PMA), A. (MO). Regional conservation status: Vulnerable (VU), Rodríguez 11586.1 (BM, INB, MO, PMA), D. Santamaría B1ab(i,iii). 5678 (BM, INB, MO, PMA), D. Santamaría 6653 (BM, INB, MO, PMA), D. Solano 4148 (BM, INB, MO, PMA), Palicourea alajuelensis C.M.Taylor. D. Solano 4803 (BM, INB, MO, PMA), E. Alfaro 5598 Terrestrial shrubs. Distribution in PILA 700–1600 m, low (BM, INB), G. Herrera 5960 (CR, INB, MO), R. Aguilar elevation mixed forest, mixed forest in transition to low 1113 (INB, MO), S. Bridgewater 4190 (BM, INB, MO, elevation forest. Global distribution: Costa Rica, Panama. PMA). Records: D. Santamaría 7598 (INB, PMA), N. Zamora 4380 (INB, PMA).

Notopleura polyphlebia (Donn.Sm.) C.M.Taylor Terrestrial herbs or shrubs. Distribution in PILA 0–1000 Palicourea albocaerulea C.M.Taylor m, low elevation mixed forest, mixed forest in transition Terrestrial shrubs or trees. Distribution in PILA 1000– to low elevation forest. Global distribution: Nicaragua to 1800 m, mixed forest in transition to low elevation Peru. Records: A.K. Monro 6270 (PMA), D. Solano 4905 forest, mixed forest in transition to cloud forest. Global (BM, INB, MO, PMA). distribution: Honduras, Costa Rica. Records: R. Aguilar 1153 (INB, MO).

Notopleura tolimensis (Wernham) C.M.Taylor Terrestrial shrubs. Distribution in PILA 800–2200 m, Palicourea angustifolia Kunth mixed forest in transition to low elevation forest, mixed Cafecillo (CR). forest in transition to cloud forest, cloud forest, low Terrestrial shrubs or trees. Distribution in PILA 800– elevation oak forest, dense thicket. Global distribution: 3200 m, mixed forest in transition to low elevation forest, Mexico to Peru. Records: A. Rodríguez 11806 (BM, INB, mixed forest in transition to cloud forest, cloud forest, MO, PMA), A. Rodríguez 11914 (BM, INB, MO, PMA), low elevation oak forest, páramo. Global distribution: D. Santamaría 4398 (INB), D. Santamaría 7042 (BM, Costa Rica to Venezuela. Records: A. Soto 1266 (INB), INB, MO, PMA), D. Solano 5001 (BM, INB, MO, PMA), C. Hamilton 3878 (MO), D. Santamaría 3768 (INB), F. Quesada 1645 (INB, MO), L. Acosta 2372 (INB), L. D. Solano 3206 (INB), D. Solano 5082 (BM, INB, MO, Angulo 160 (INB), L. Angulo 169 (INB), L. Angulo 318 PMA), G. Davidse 23782 (CR, MO), L. González 3622 (INB), M. Alfaro 21 (INB). (INB), L. González 3665 (INB), M. Alfaro 137 (INB), N. Zamora 4093 (BM, INB, MO, PMA), N. Zamora 4105

Notopleura tonduzii (Standl.) C.M.Taylor (BM, INB, MO, PMA) , N. Zamora 4126 (BM, INB, MO, Terrestrial shrubs. Distribution in PILA 300–1500 m, low PMA). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Palicourea calophlebioides C.M.Taylor Global distribution: Nicaragua to Panama. Records: D. Terrestrial shrubs. Distribution in PILA 1800–2700 m, Solano 4188 (BM, INB, MO, PMA). cloud forest, low elevation oak forest, high elevation oak forest. Global distribution: Endemic to Costa Rica.

Notopleura uliginosa (Sw.) Bremek. Records : A. Rodríguez 10583 (BM, INB, MO, PMA), Terrestrial herbs. Distribution in PILA 800–1800 m, D. Solano 3956 (BM, INB, MO, PMA), D. Solano 4061 mixed forest in transition to low elevation forest, mixed (BM, INB, MO, PMA), G. Herrera 5952 (CR, INB, MO), forest in transition to cloud forest, dense thicket. Global G. Herrera 5994 (CR, INB, MO). distribution: Mexico to the Guyanas, Bolivia, Greater Antilles, Tri. Records: A.K. Monro 6314 (PMA), D. Palicourea chiriquina Standl. Santamaría 6747 (BM, INB, MO, PMA), D. Solano Terrestrial shrubs. Distribution in PILA 1400–2300 m, 4840 (BM, INB, MO, PMA), D. Solano 5015 (BM, INB, mixed forest in transition to cloud forest, cloud forest, MO, PMA), D. Solano 5028 (BM, INB, MO, PMA), L. low elevation oak forest. Global distribution: Costa Rica, Martínez 335 (PMA). Panama. Records: A. Fernández 685 (CR, INB, MO), A. Fernández 732 (CR, INB, MO), A. Rodríguez 10858

ψ Palicourea adusta Standl. (BM, INB, MO, PMA) , G. Herrera 6263 (CR, INB, MO), Cafecillo, Candelilla (CR). J. Bittner 1786 (INB, MO), M. Grayum 10342 (INB). Terrestrial shrubs. Distribution in PILA 1500–3200

m, mixed forest in transition to cloud forest, cloud forest, Palicourea discolor K.Krause Cafecillo (CR) .

Terrestrial shrubs or trees. Distribution in PILA G. Davidse 25192 (MO), G. Davidse 28857 (CR, MO), 1200–2800 m, mixed forest in transition to low elevation G. Davidse 28933 (CR, MO), G. Davidse s.n. (CR), G. forest, mixed forest in transition to cloud forest, cloud Herrera 5201 (CR, INB, MO), G. Herrera 5463 (INB), forest, low elevation oak forest, high elevation oak forest. G. McPherson 15920 (MO), J. Bittner 1743 (INB, MO), Global distribution: Costa Rica, Panama. Records: E. J. González 793 (CR, INB, MO). Alfaro 2264 (CR, INB), G. Davidse 28430 (CR, MO), J. González 796 (CR, INB, MO), L. Angulo 316 (INB), M. Palicourea matamana C.M.Taylor Moraga 571 (INB, MO). Terrestrial shrubs. Distribution in PILA 1400– 1400 m, mixed forest in transition to cloud forest. Global

Palicourea garciae Standl. distribution: Costa Rica, Panama. Records: G. Herrera Cafecillo (CR) . 2805 (MO). Terrestrial shrubs or trees. Distribution in PILA 1100– 2500 m, mixed forest in transition to low elevation forest, Palicourea montivaga Standl. mixed forest in transition to cloud forest, cloud forest, low Terrestrial shrubs. Distribution in PILA 1400–3100 m, elevation oak forest. Global distribution: Costa Rica to mixed forest in transition to cloud forest, cloud forest, low Peru. Records: A.K. Monro 4478 (BM, MO), A.K. Monro elevation oak forest, high elevation oak forest, páramo. 4527 (BM, MO), A.K. Monro 5596 (BM, INB, MO, Global distribution: Costa Rica, Panama. Records: A. PMA). Rodríguez 10697 (BM, INB, MO, PMA), A. Rodríguez 10730 (BM, INB, MO, PMA), A. Soto 300 (INB), D.

Palicourea gomezii C.M.Taylor Solano 3937 (BM, INB, MO, PMA), E. Alfaro 3226 Terrestrial shrubs. Distribution in PILA 500–1500 m, low (INB), E. Alfaro 3248 (INB), E. Alfaro 3944 (INB), E. elevation mixed forest, mixed forest in transition to low Alfaro 4616 (INB), J. F. Morales 5798 (INB), F. Quesada elevation forest, mixed forest in transition to cloud forest. 1606 (INB, MO), F. Quesada 1823 (INB), F. Quesada Global distribution: Costa Rica to Peru. Records: A. 1922 (INB), L. Angulo 205 (INB), M. Moraga 378 (INB), Rodríguez 11589 (BM, INB, MO, PMA), D. Santamaría S. Knapp 9923 (BM, INB, MO). 6668 (BM, INB, MO, PMA), D. Solano 4899 (BM, INB, MO, PMA), G. Herrera 2840 (MO). Palicourea nebulosa (Dwyer) C.M.Taylor Terrestrial trees. Distribution in PILA 300–2000 m, low

Palicourea hammelii C.M.Taylor elevation mixed forest, mixed forest in transition to low Terrestrial shrubs. Distribution in PILA 1000–2200 m, elevation forest, mixed forest in transition to cloud forest. mixed forest in transition to low elevation forest, mixed Global distribution: Endemic to Panama. Records: von forest in transition to cloud forest, cloud forest, low Hagen 2156 (NY). elevation oak forest. Global distribution: Costa Rica, Panama. Records: G. Herrera 2781 (CR, MO). Palicourea orosiana C.M.Taylor Terrestrial shrubs. Distribution in PILA 1200–2000 m,

Palicourea hondensis (Standl.) C.M.Taylor mixed forest in transition to low elevation forest, mixed Terrestrial shrubs or trees. Distribution in PILA 0–1300 forest in transition to cloud forest. Global distribution: m, low elevation mixed forest, mixed forest in transition Costa Rica, Panama. Records: A. Fernández 1002 (INB), to low elevation forest, mixed forest in transition to A. Fernández 1006 (INB), E. Alfaro 4710 (INB). cloud forest. Global distribution: Costa Rica, Panama. Records: A. Rodríguez 11553,1 (BM, INB, MO, PMA), Palicourea padifolia (Humb. & Bonpl. ex Schult.) D. Santamaría 6493 (BM, INB, MO, PMA), D. Solano C.M.Taylor & Lorence 4841 (BM, INB, MO, PMA), G. Herrera 3128 (MO). Cafecillo (CR). Terrestrial shrubs or trees. Distribution in PILA

Palicourea lasiorrhachis Oerst. 600–2500 m, low elevation mixed forest, mixed forest Zorrillo amarillo, Cafecillo, Zorrillo (CR) . in transition to low elevation forest, mixed forest in Terrestrial shrubs. Distribution in PILA 700–3000 transition to cloud forest, cloud forest, low elevation oak

m, low elevation mixed forest, mixed forest in transition forest. Global distribution: Mexico to Peru. Records: A.K. to low elevation forest, mixed forest in transition to Monro 4890 (BM, INB, MO), A. Rodríguez 11733 (BM, cloud forest, cloud forest, low elevation oak forest, high INB, MO, PMA), C. Hamilton 3601 (MO), C. Hamilton elevation oak forest. Global distribution: Costa Rica to 3876 (MO), C. Hamilton 777 (MO), D. Santamaría 5921 Ecuador. Records: A. Chacón 254 (CR, INB, MO), A.K. (BM, INB, MO, PMA), D. Santamaría 7268 (BM, INB, Monro 4414 (BM, INB, MO), A.K. Monro 4934 (BM, MO, PMA), D. Solano 2896 (INB), D. Solano 2925 (INB), INB, MO), D. Santamaría 5872 (BM, INB, MO, PMA), D. Solano 5201 (BM, INB, MO, PMA), E. Alfaro 3194 D. Solano 3882 (BM, INB, MO, PMA), D. Solano 3966 (INB), E. Mora 296 (CR, INB), G. Davidse 23778 (CR, (BM, INB, MO, PMA), G. Davidse 25159 (CR, MO), MO), G. Herrera 2715 (MO), J. Bittner 1441 (MO), J. González 795 (CR, INB, MO), L. Angulo 170 (CR, INB), INB, MO, PMA), D. Santamaría 6637 (BM, INB, MO, L. González 3643 (INB), N. Zamora 4100 (BM, INB, PMA), D. Solano 4809 (BM, INB, MO, PMA), E. Alfaro MO, PMA), N. Zamora 4172 (BM, INB, MO, PMA), N. 5590 (BM, MO), E. Navarro 681 (INB). Zamora 4207 (BM, INB, MO, PMA), R. Villalobos 250 (CR, INB). Palicourea triphylla DC. Terrestrial herbs or shrubs. Distribution in PILA 100–1300

Palicourea pauciflora Standl. m, low elevation mixed forest, mixed forest in transition Terrestrial shrubs. Distribution in PILA 1100–2200 m, to low elevation forest, mixed forest in transition to cloud mixed forest in transition to low elevation forest, mixed forest. Global distribution: Mexico, Guatemala to the forest in transition to cloud forest, cloud forest, low Guyanas, Bolivia, Greater Antilles. Records: D. Solano elevation oak forest. Global distribution: Costa Rica, 5064 (BM, INB, MO, PMA). Panama. Records: G. Herrera 5452 (CR, INB, MO). Palicourea umbelliformis (Dwyer & M.V.Hayden)

Palicourea purpurea C.M.Taylor C.M.Taylor Terrestrial shrubs or trees. Distribution in PILA 800–3300 Terrestrial shrubs or trees. Distribution in PILA 0–1400 m, mixed forest in transition to low elevation forest, m, low elevation mixed forest, mixed forest in transition mixed forest in transition to cloud forest, cloud forest, low to low elevation forest, mixed forest in transition to cloud elevation oak forest, páramo. Global distribution: Costa forest. Global distribution: Costa Rica, Panama. Records: Rica to Colombia. Records: A.K. Monro 4235 (BM, INB, A.K. Monro 4281 (BM, INB, MO), D. Santamaría 7682 MO), A.K. Monro 4318 (BM, INB, MO), A.K. Monro (INB, PMA). 4405 (BM, INB, MO), A.K. Monro 5502 (BM, INB, MO, PMA), A.K. Monro 5534 (BM, INB, MO, PMA), Palicourea vestita Standl. A.K. Monro 5577 (BM, INB, MO, PMA), A. Rodríguez Terrestrial shrubs or trees. Distribution in PILA 1100– 10992 (BM, INB, MO, PMA), D. Solano 3880 (BM, INB, 2500 m, mixed forest in transition to low elevation forest, MO, PMA), D. Solano 3959 (BM, INB, MO, PMA), G. mixed forest in transition to cloud forest, cloud forest, Davidse 29073 (CR, MO), G. McPherson 15911 (MO). low elevation oak forest. Global distribution: Costa Rica, Panama. Records: A. Rodríguez 10687 (BM, INB, MO,

ψ Palicourea salicifolia Standl. PMA ), A. Rodríguez 9862 (INB), A. Soto 1267 (INB), Terrestrial shrubs. Distribution in PILA 1200–3100 m, D. Santamaría 5694 (BM, INB, MO, PMA), D. Solano mixed forest in transition to low elevation forest, mixed 3904 (BM, INB, MO, PMA), G. Herrera 6082 (CR, INB, forest in transition to cloud forest, cloud forest, low MO), G. Herrera 6129 (CR, INB, MO), L. González 3621 elevation oak forest, high elevation oak forest, páramo. (INB). Global distribution: Costa Rica, Panama. Records: A. Rodríguez 10704 (BM, INB, MO, PMA), D. Santamaría Pentagonia costaricensis (Standl.) W.C.Burger & 6666 (BM, INB, MO, PMA), D. Solano 5087 (BM, INB, C.M.Taylor MO, PMA), D. Solano 5133 (BM, INB, MO, PMA), E. Terrestrial trees. Distribution in PILA 0–2200 m, low Alfaro 3650 (INB), L.Angulo 262 (INB), L. González 3624 elevation mixed forest, mixed forest in transition to low (INB), N. Zamora 4094 (BM, INB, MO, PMA). Global elevation forest, mixed forest in transition to cloud forest, conservation status: Endangered (EN), B1ab(i,iii). cloud forest, low elevation oak forest. Global distribution: Nicaragua to Colombia. Records: A. Rodríguez 11815

Palicourea skotakii C.M.Taylor (BM, INB, MO, PMA) , D. Santamaría 7028 (BM, INB, Terrestrial shrubs. Distribution in PILA 2300–2900 MO, PMA), D. Solano 4428 (BM, INB, MO, PMA), D. m, low elevation oak forest, high elevation oak forest. Solano 5026 (BM, INB, MO, PMA). Global distribution: Costa Rica to Colombia. Records: B. Gamboa 1213 (INB, MO), D. Solano 4052 (BM, INB, •, □ Pentagonia gomez-lauritoi Hammel MO, PMA ), E. Alfaro 3472 (INB), E. Alfaro 4656 (INB), Terrestrial shrubs. Distribution in PILA 0–800 m, low G. Davidse 25125 (CR, MO), G. Davidse 28879 (CR, elevation mixed forest, mixed forest in transition to low MO), G. Davidse 29004 (MO), L. Angulo 237 (INB), L. elevation forest. Global distribution: Costa Rica, Panama. González 3615 (INB). Records: L. Gómez 24065 (MO), G. Herrera 3092 (MO).