Palicourea standleyana C.M.Taylor

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Palicourea standleyana C.M.Taylor Posoqueria costaricensis C.M.Taylor

Terrestrial shrubs or trees. Distribution in PILA 1100– Boca de vieja, Cacica, Fruta de mono, Guayaba de mico, 2400 m, mixed forest in transition to low elevation forest, Guayabillo de montaña, Wolf coffe, Guayaba de mono mixed forest in transition to cloud forest, cloud forest, low ( CR), Boca de vieja, Borojó, Fruta de mono, Tuliviejo elevation oak forest, pasture. Global distribution: Costa (P).

Rica to Ecuador. Records: D. Santamaría 5877 ( BM, Terrestrial trees. Distribution in PILA 500–1700 m,

low elevation mixed forest, mixed forest in transition to Terrestrial shrubs. Distribution in PILA 200–1400 m, low low elevation forest, mixed forest in transition to cloud elevation mixed forest, mixed forest in transition to low forest, dense thicket. Global distribution: Mexico to elevation forest, mixed forest in transition to cloud forest. Brazil. Records: E. Alfaro 2198 ( INB) . Global distribution: Endemic to Costa Rica. Records: A. Jiménez 868 ( MO) , F. Solís 32 ( INB) .

Posoqueria grandifructa Hammel & C.M.Taylor Terrestrial shrubs or trees. Distribution in PILA 0–1200 Psychotria berteroana DC. m, low elevation mixed forest, mixed forest in transition Terrestrial shrubs or trees. Distribution in PILA 0–2200 to low elevation forest. Global distribution: Mexico, m, low elevation mixed forest, mixed forest in transition Guatemala, Costa Rica to Venezuela. Records: G. Herrera to low elevation forest, mixed forest in transition to cloud 2711 (MO). forest, cloud forest, low elevation oak forest. Global distribution: Mexico to Honduras, Costa Rica, Panama,

Psychotria acuminata Benth. Ecuador, Peru, Greater Antilles, Lesser Antilles. Records: Terrestrial shrubs. Distribution in PILA 0–1400 m, low A. Rodríguez 11769 ( BM, INB, MO, PMA). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Psychotria buchtienii (H.J.P.Winkl.) Standl. Global distribution: Mexico to Peru, Greater Antilles, Terrestrial shrubs. Distribution in PILA 0–1500 m, low Lesser Antilles. Records: D. Solano 5807 ( BM, INB, elevation mixed forest, mixed forest in transition to MO). low elevation forest, mixed forest in transition to cloud forest. Global distribution: Mexico to Venezuela, Bolivia.

ψ Psychotria angustiflora K.Krause Records : A. Chacón 298 (CR, MO), D. Santamaría 5941 Terrestrial shrubs or trees. Distribution in PILA 100–1600 (BM, INB, MO, PMA), D. Santamaría 6491 (BM, INB, m, low elevation mixed forest, mixed forest in transition MO, PMA). to low elevation forest. Global distribution: Costa Rica to Colombia. Records: D. Santamaría 6206 (BM, INB, MO, Psychotria calophylla Standl. PMA ), D. Solano 4356 (BM, INB, MO, PMA), D. Solano Terrestrial shrubs. Distribution in PILA 500–1800 m, low 4459 (BM, INB, MO, PMA), S. Bridgewater 4155 (BM, elevation mixed forest, mixed forest in transition to low INB, MO, PMA). Regional conservation status: Near elevation forest, mixed forest in transition to cloud forest. Threatened (NT). Global distribution: Mexico, Guatemala, Panama to Peru. Records: C. Hamilton 776 (MO).

Psychotria aubletiana Steyerm. Cafecillo (CR) . Psychotria chiriquensis (Standl.) C.M.Taylor Terrestrial herbs or shrubs. Distribution in PILA 800– Terrestrial shrubs. Distribution in PILA 1200–3000 m, 2700 m, mixed forest in transition to low elevation forest, mixed forest in transition to low elevation forest, mixed mixed forest in transition to cloud forest, cloud forest, forest in transition to cloud forest, cloud forest, low low elevation oak forest, high elevation oak forest, dense elevation oak forest, high elevation oak forest, dense thicket. Global distribution: Mexico to the Guyanas, thicket. Global distribution: Mexico, Costa Rica, Panama. Lesser Antilles, Greater Antilles. Records: A. Fernández Records: A. Fernández 888 (CR, INB, MO), A.K. Monro 877 (CR, INB, MO), A.K. Monro 4233 (BM, MO), A.K. 5474 (BM, INB, MO, PMA), A.K. Monro 5624 (BM, Monro 4845 (BM, MO), A.K. Monro 5059 (BM, INB), INB, MO, PMA), A. Rodríguez 10647 (BM, INB, MO, A.K. Monro 5532 (BM, INB, MO, PMA), A.K. Monro PMA), A. Rodríguez 11496 (BM, INB, MO, PMA), D. 5593 (BM, INB, MO, PMA), A.K. Monro 5630 (BM, INB, Santamaría 6626 (BM, INB, MO, PMA), D. Solano 3847 MO, PMA), A. Rodríguez 10665 (BM, INB, MO, PMA), (BM, INB, MO, PMA), D. Solano 4661 (BM, INB, MO, A. Rodríguez 11782 (BM, INB, MO, PMA), A. Soto 1264 PMA), G. Davidse 29131 (CR, MO). (INB), D. Santamaría 3757 (INB), D. Santamaría 5687 (BM, INB, MO, PMA), D. Santamaría 6643 (BM, INB, Psychotria chiriquina Standl. MO, PMA ), D. Solano 3204 (INB), D. Solano 3874 (BM, Terrestrial shrubs or trees. Distribution in PILA 1400– INB, MO, PMA), D. Solano 4664 (BM, INB, MO, PMA), 3000 m, mixed forest in transition to cloud forest, cloud D. Solano 4779 (BM, INB, MO, PMA), D. Solano 5081 forest, low elevation oak forest, high elevation oak forest. (BM, INB, MO, PMA), G. Davidse 28379 (CR, MO), G. Global distribution: Nicaragua to Colombia. Records: G. Davidse 28684 (CR, MO), G. McPherson 15880 (MO), J. McPherson 15899 (MO), M.A. Thomas 1615 (MO). Bittner 1477 (INB, MO), J. Bittner 1526 (INB, MO), J. Bittner 1837 (INB), L. Angulo 239 (INB), M. Valerio 188 Psychotria convergens C.M.Taylor (CR, MO) , S. Bridgewater 4193 (BM, INB, MO, PMA). Terrestrial shrubs. Distribution in PILA 1100–2600 m, mixed forest in transition to low elevation forest, mixed

Psychotria aurantiibractea C.M.Taylor forest in transition to cloud forest, cloud forest, low elevation oak forest, high elevation oak forest. Global Fernández 1004 (CR, INB), A.K. Monro 4360 (BM, INB, distribution: Costa Rica to Peru. Records: A.K. Monro MO), A.K. Monro 4939 (BM, INB, MO), A.K. Monro 4499 (BM, MO), B. Gamboa 1220 (CR, INB, MO), G. 4949 (BM, INB, MO), A. Rodríguez 11323 (BM, INB, Davidse 25751 (CR, MO), G. Davidse 28628 (CR, MO), MO, PMA), G. Herrera 3280 (INB, MO), G. Herrera G. Davidse 29227 (MO). 5262 (MO), G. Herrera 6042 (CR, INB, MO), J. Bittner 1528 (INB, MO), R. Aguilar 1136 (INB, MO).

Psychotria croceovenosa Dwyer Terrestrial shrubs. Distribution in PILA 1200–1700 m, ψ Psychotria graciliflora Benth. mixed forest in transition to low elevation forest, mixed Terrestrial shrubs. Distribution in PILA 0–2000 m, low forest in transition to cloud forest. Global distribution: elevation mixed forest, mixed forest in transition to low Costa Rica, Panama. Records: D. Santamaría 6636 (INB, elevation forest, mixed forest in transition to cloud forest, MO). cloud forest, dense thicket. Global distribution: Mexico to Colombia. Records: A.K. Monro 4294 (BM, INB, MO),

Psychotria cyanococca Seem. ex Dombrain A.K. Monro 4724 (BM, INB, MO), D. Solano 5011 (BM, Terrestrial shrubs. Distribution in PILA 0–1400 m, low INB, MO, PMA). Regional conservation status: Near elevation mixed forest, mixed forest in transition to low Threatened (NT). elevation forest, mixed forest in transition to cloud forest. Global distribution: Mexico, Guatemala, Nicaragua to Psychotria guapilensis (Standl.) Hammel Ecuador. Records: D. Santamaría 7688 (PMA), D. Solano Terrestrial herbs to shrubs. Distribution in PILA 0–1200 4275 (BM, INB, MO, PMA), D. Solano 4308 (BM, INB, m, low elevation mixed forest, mixed forest in transition to MO, PMA). low elevation forest, secondary forest. Global distribution: Nicaragua to Ecuador. Records: A.K. Monro 6287 (PMA),

Psychotria dichroa (Standl.) C.M.Taylor S. Bridgewater 4170 (BM, INB, MO, PMA). Cafecillo (CR). Terrestrial shrubs. Distribution in PILA 1100–2100 Psychotria hazenii Standl.

m, mixed forest in transition to low elevation forest, Terrestrial shrubs. Distribution in PILA 1100–1600 m, mixed forest in transition to cloud forest, cloud forest, mixed forest in transition to low elevation forest. Global low elevation oak forest. Global distribution: Costa Rica, distribution: Costa Rica to Ecuador. Records: D. Solano Panama. Records: A. De Sedas 564 (PMA), A. Fernández 4547 (BM, INB, MO, PMA). 1023 (INB), A.K. Monro 4935 (BM, INB, MO), A.K. Monro 6036 (PMA), A.K. Monro 6212 (PMA), D. □ Psychotria jimenezii Standl. Santamaría 7733 (PMA). Terrestrial shrubs or trees. Distribution in PILA 1100– 1700 m, mixed forest in transition to low elevation

ψ Psychotria elata (Sw.) Hammel forest, mixed forest in transition to cloud forest. Global Botón rojo, Manga de señora, Labios de mujer, Labios de distribution: Costa Rica, Panama. Records: A. Rodríguez puta, Labios de novia (CR). 11303 (BM, INB, MO, PMA), D. Solano 4351 (BM, INB, Terrestrial shrubs or trees.Distribution in PILA0–3800 MO, PMA), D. Solano 4352 (BM, INB, MO, PMA), D.

m, low elevation mixed forest, mixed forest in transition Solano 4884 (BM, INB, MO, PMA). to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo, Psychotria marginata Sw. pasture, dense thicket. Global distribution: Mexico to (CR), Cafecillo (P). Ecuador, Greater Antilles, Lesser Antilles. Records: A. Terrestrial shrubs or trees. Distribution in PILA Fernández 1005 (CR, INB), A.K. Monro 4806 (BM, INB, 0–1000 m, low elevation mixed forest, mixed forest in MO), D. Santamaría 6480 (BM, INB, MO, PMA), D. transition to low elevation forest,, dense thicket. Global Santamaría 6689 (BM, INB, MO, PMA), D. Santamaría distribution: Mexico to Bolivia Antilles. Records: A.K. 7343 (PMA), D. Santamaría 7756 (PMA), E. Alfaro Monro 5084 (BM, INB). 5538 (BM, INB, MO), G. Gallardo 98 (CR, INB, MO), S. Bridgewater 4185 (BM, INB, MO, PMA). Regional Psychotria microbotrys Ruiz ex Standl. conservation status: Near Threatened (NT). Terrestrial shrubs or trees. Distribution in PILA 0–2600 m, low elevation mixed forest, mixed forest in transition

Psychotria goldmanii Standl. to low elevation forest, mixed forest in transition to Cafecillo (CR). cloud forest, cloud forest, low elevation oak forest, high Terrestrial shrubs or trees. Distribution in PILA elevation oak forest. Global distribution: Nicaragua to 1200–2000 m, mixed forest in transition to low elevation Peru. Records: A.K. Monro 5666 (BM, INB, MO, PMA), forest, mixed forest in transition to cloud forest. Global A.K. Monro 6148 (PMA), D. Santamaría 5966 (BM, distribution: Costa Rica to Colombia. Records: A. INB, MO, PMA), D. Santamaría 6466 (BM, INB, MO, PMA). Terrestrial shrubs. Distribution in PILA 1000–2700 m, mixed forest in transition to low elevation forest, mixed

Psychotria nubiphila Dwyer forest in transition to cloud forest, cloud forest, low Terrestrial shrubs or trees. Distribution in PILA 900–2300 elevation oak forest, high elevation oak forest. Global m, mixed forest in transition to low elevation forest, distribution: Costa Rica to Colombia. Records: G. Davidse mixed forest in transition to cloud forest, cloud forest, 25165 (CR, MO), G. Davidse 29061 (CR, MO). low elevation oak forest. Global distribution: Mexico, Costa Rica, Panama. Records: A.K. Monro 4352 (BM), Psychotria psychotriifolia (Seem.) Standl. C. Hamilton 763 (MO), G. Davidse 28389 (CR, MO), L. Terrestrial shrubs. Distribution in PILA 0–800 m, low Acosta 2336 (INB), L. Donzo 2 (INB). elevation mixed forest, mixed forest in transition to low elevation forest. Global distribution: Nicaragua to

Psychotria orosiana Standl. Ecuador, Venezuela. Records: D. Santamaría 6164 (BM, Terrestrial shrubs. Distribution in PILA 600–1800 m, low INB, MO, PMA). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, Psychotria recordiana Standl. pasture. Global distribution: Costa Rica to Colombia. Terrestrial shrubs. Distribution in PILA 900–2200 m, Records: D. Santamaría 5917 (BM, INB, MO, PMA). mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low

Psychotria panamensis Standl. elevation oak forest. Global distribution: Honduras to Terrestrial shrubs or trees. Distribution in PILA 0–2000 Ecuador, Venezuela. Records: B. Gamboa 629 (INB), D. m, low elevation mixed forest, mixed forest in transition Santamaría 6202 (BM, INB, MO, PMA), E. Alfaro 3191 to low elevation forest, mixed forest in transition to (INB), G. Davidse 23775 (CR, MO), L. Acosta 2395 cloud forest, cloud forest. Global distribution: Mexico to (INB), A. Rodríguez 11201 (BM, INB, MO, PMA), A. Colombia. Records: L. Acosta 2405 (INB). Rodríguez 11795 (BM, INB, MO, PMA), D. Solano 2912 (INB).

Psychotria paradichroa C.M.Taylor Terrestrial shrubs. Distribution in PILA 1500–1700 Psychotria saltatrix C.M.Taylor m, mixed forest in transition to cloud forest. Global Shrubs. Distribution in PILA 1200–1700 m, mixed distribution: Endemic to Panama. Records: A.K. Monro forest in transition to low elevation forest, mixed forest 4223 (BM, MO). in transition to cloud forest. Global distribution: Costa Rica to Ecuador. Records: A. Fernández 1018 (CR, INB,

Psychotria parvifolia Benth. MO ). Cafecillo (CR). Terrestrial shrubs. Distribution in PILA 1000–2200 Psychotria sarapiquensis Standl.

m, mixed forest in transition to low elevation forest, Terrestrial shrubs or trees. Distribution in PILA 600–1600 mixed forest in transition to cloud forest, cloud forest, m, low elevation mixed forest, mixed forest in transition low elevation oak forest. Global distribution: Guatemala, to low elevation forest. Global distribution: Mexico to Costa Rica, Panama. Records: A.K. Monro 5039 (BM, Ecuador. Records: A. Rodríguez 11287 (BM, INB, MO, INB, MO), G. Davidse 23786 (CR, MO). PMA), D. Solano 4413 (BM, INB, MO, PMA).

Psychotria pilosa Ruiz & Pav. Psychotria sixaolensis C.W.Ham. Terrestrial shrubs. Distribution in PILA 0–2300 m, low Terrestrial shrubs. Distribution in PILA 0–700 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest. Global distribution: Costa Rica, elevation forest, mixed forest in transition to cloud forest, Panama. Records: G. Herrera 3077 (MO). cloud forest, low elevation oak forest. Global distribution: Nicaragua to Venezuela, Bolivia. Records: A. De Sedas Psychotria sylvivaga Standl. 578 (PMA), A.K. Monro 4256 (BM, INB, MO), A.K. Terrestrial shrubs. Distribution in PILA 1400–3200 m, Monro 5674 (BM, INB, MO, PMA), D. Santamaría 6826 mixed forest in transition to cloud forest, cloud forest, (BM, INB, MO, PMA), D. Santamaría 7583 (PMA), D. low elevation oak forest, páramo. Global distribution: Santamaría 7636 (PMA), D. Santamaría 7666 (PMA), Costa Rica to Ecuador. Records: E. Alfaro 3484 (INB), E. D. Solano 4346 (BM, INB, MO, PMA), D. Solano 4401 Alfaro 3626 (INB), E. Alfaro 3985 (INB), E. Alfaro 4599 (BM, INB, MO, PMA), D. Solano 4848 (BM, INB, MO, (INB), G. Davidse 25711 (CR, MO), G. Herrera 3649 PMA). (INB), G. Herrera 6023 (INB, MO), J. F. Morales 5799 (INB, MO), F. Quesada 1604 (INB), F. Quesada 1904

Psychotria pisonioides Standl. (INB) , F. Quesada 1906 (INB), L. Angulo 206 (INB), L. Angulo 231 (INB), M. Grayum 10371 (CR, INB, MO), N.

Zamora 2711 (INB), N. Zamora 2714 (INB), N. Zamora Costa Rica, Panama. Records: D. Santamaría 5944 4170 (BM, INB, MO, PMA), A. Rodríguez 11872 (BM, (INB). INB, MO, PMA), A. Soto 299 (INB), D. Santamaría 7227 (BM, INB, MO, PMA). Raritebe palicoureoides Wernham Terrestrial shrubs or trees. Distribution in PILA 200–1700

Psychotria torresiana Standl. m, low elevation mixed forest, mixed forest in transition Terrestrial shrubs. Distribution in PILA 1000–1800 m, to low elevation forest, mixed forest in transition to cloud mixed forest in transition to low elevation forest, mixed forest, pasture, secondary forest. Global distribution: forest in transition to cloud forest. Global distribution: Costa Rica to Colombia. Records: A.K. Monro 5781 (BM, Costa Rica, Panama. Records: J. Bittner 1528 (INB, MO) INB, MO, PMA), A.K. Monro 6322 (PMA), A. Rodríguez Aguilar 1136 (INB, MO), A. De Sedas 561 (INB, PMA), 11409 (BM, INB, MO, PMA), D. Santamaría 5928 (BM, Fernández 1004 (CR, INB), Herrera 5262 (INB, MO), INB, MO, PMA), D. Santamaría 6484 (BM, INB, MO, A.K. Monro 4360 (BM, INB, MO), D. Santamaría 7339 PMA), D. Santamaría 6525 (BM, INB, MO, PMA), D. (INB, PMA), D. Solano 5375 (INB, PMA). Santamaría 6691 (BM, INB, MO, PMA), D. Santamaría 6738 (BM, INB, MO, PMA), D. Santamaría 7352 (PMA),

Psychotria trichotoma M.Martens & Galeotti D. Santamaría 7734 (PMA), D. Solano 4659 (BM, INB, Terrestrial shrubs. Distribution in PILA 100–1500 m, low MO, PMA), D. Solano 4743 (BM, INB, MO, PMA), D. elevation mixed forest, mixed forest in transition to low Solano 5443 (PMA), E. Alfaro 5547 (BM, INB, MO), G. elevation forest, mixed forest in transition to cloud forest. Herrera 6203 (INB), L. Martínez 353 (PMA). Global distribution: Mexico to Bolivia. Records: M.A. Thomas 1586 (MO). ψ Rogiera amoena Planch. Quina, Teresa (CR) .

Psychotria tsakiana C.M.Taylor Terrestrial shrubs or trees. Distribution in PILA Terrestrial shrubs. Distribution in PILA 0–500 m, low 600–3400 m, low elevation mixed forest, mixed forest elevation mixed forest. Global distribution: Costa Rica to in transition to low elevation forest, mixed forest in Bolivia. Records: G. Herrera 3035 (MO). transition to cloud forest, cloud forest, low elevation oak forest, páramo. Global distribution: Mexico to Colombia.

Psychotria viridis Ruiz & Pav. Records : B. Gamboa 584 (CR, INB, MO), J. González Terrestrial shrubs. Distribution in PILA 0–1300 m, low 3630 (INB), C. Hamilton 3637 (MO), C. Hamilton 3905 elevation mixed forest, mixed forest in transition to low (MO), A.K. Monro 4668 (BM, INB, MO), A.K. Monro elevation forest, mixed forest in transition to cloud forest. 5951 (PMA), D. Solano 5231 (BM, INB, MO, PMA), N. Global distribution: Mexico to Bolivia, Greater Antilles. Zamora 4088 (BM, INB, MO, PMA), N. Zamora 4131 Records: A. Rodríguez 11198 (BM, INB, MO, PMA), D. (BM, INB, MO, PMA). Regional conservation status: Santamaría 7679 (PMA), D. Solano 4295 (BM, INB, Near Threatened (NT). MO, PMA). Ronabea latifolia Aubl.

Randia calycosa Standl. Terrestrial shrubs. Distribution in PILA 0–1500 m, low Terrestrial shrubs or trees.Distribution in PILA 1000–1900 elevation mixed forest, mixed forest in transition to low m, mixed forest in transition to low elevation forest, mixed elevation forest, mixed forest in transition to cloud forest. forest in transition to cloud forest. Global distribution: Global distribution: Mexico to Honduras to Bolivia. Costa Rica, Panama. Records: D. Solano 5207 (BM, INB, Records: A. Chacón 188 (CR, INB, MO). MO, PMA), F. Quesada 1678 (INB, MO), L. Angulo 140 (INB, MO), M. Moraga 265 (INB, MO). Rudgea laevis C.M.Taylor Terrestrial shrubs. Distribution in PILA 1000–1700 m,

Randia gentryi Dwyer mixed forest in transition to low elevation forest, mixed Terrestrial trees. Distribution in PILA 100–1400 m, low forest in transition to cloud forest. Global distribution: elevation mixed forest, mixed forest in transition to low Endemic to Costa Rica. Records: A. Rodríguez 11866 elevation forest, mixed forest in transition to cloud forest. (INB, CR), D. Solano 5066 (INB). Global distribution: Costa Rica to Colombia. Records: S. Bridgewater 4160 (BM, INB, MO, PMA). Rudgea skutchii Standl. Terrestrial shrubs or trees. Distribution in PILA 0–1200

Randia vazquezii Lorence & Dwyer m, low elevation mixed forest, mixed forest in transition Terrestrial shrubs. Distribution in PILA 700–900 m, low to low elevation forest. Global distribution: Costa Rica elevation mixed forest, mixed forest in transition to low to Peru. Records: A.K. Monro 6216 (PMA), A.K. Monro elevation forest. Global distribution: Mexico, Guatemala, 6277 (PMA), L. Martínez 357 (PMA).

Rudgea trifurcata Gómez-Laur. Terrestrial herbs. Distribution in PILA 0–1800 m, low Terrestrial shrubs or trees. Distribution in PILA 0–500 elevation mixed forest, mixed forest in transition to low m, low elevation mixed forest. Global distribution: Costa elevation forest, mixed forest in transition to cloud forest, Rica, Panama. Records: A. Rodríguez 11206 (BM, INB, cloud forest. Global distribution: Mexico to Paraguay, MO, PMA), D. Solano 4363 (BM, INB, MO, PMA), D. Caribbean. Records: D. Solano 2840 (INB), D. Solano Solano 4392 (BM, INB, MO, PMA). 4911 (BM, INB, MO, PMA), M. Alfaro 8 (INB).

Sabicea panamensis Wernham Spermacoce remota Lam. Terrestrial twiners. Distribution in PILA 0–1700 m, Terrestrial herbs. Distribution in PILA 0–2400 m, low low elevation mixed forest, mixed forest in transition to elevation mixed forest, mixed forest in transition to low low elevation forest, mixed forest in transition to cloud elevation forest, mixed forest in transition to cloud forest, forest, cloud forest. Global distribution: Mexico to Peru. cloud forest, low elevation oak forest. Global distribution: Records: A.K. Monro 5801 (BM, INB, MO, PMA), A.K. Mexico to Ecuador, Greater Antilles, Africa, Madagascar. Monro 6361 (PMA), A. Rodríguez 11605 (BM, INB, MO, Records: R. J. Schmalzel 37 (MO). PMA), D. Santamaría 6455 (BM, INB, MO, PMA), L. Martínez 391 (PMA). Spermacoce remota Lam. Botoncillo, Verdolaga (CR) .

Sabicea villosa Willd. ex Schult. Terrestrial herbs. Distribution in PILA 0–2400 m, low Terrestrial twiners. Distribution in PILA 100–900 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, elevation forest. Global distribution: Mexico to Bolivia. cloud forest, low elevation oak forest. Global distribution: Records: D. Solano, Oct. 25, 2007 (Field obs.) . El Salvador, Costa Rica to Peru, French Guyana, the Guyanas , Greater Antilles, Lesser Antilles. Records: C .

Schradera neeoides Standl. ex Steyerm. Hamilton 703 (MO), M.A. Thomas 1647 (MO). Epiphytic shrubs. Distribution in PILA 1000–1500 m, mixed forest in transition to low elevation forest, mixed RUTACEAE forest in transition to cloud forest. Global distribution: Costa Rica to Ecuador. Records: L. Martínez 389 (BM, Amyris brenesii Standl. INB, MO, PMA ). Terrestrial shrubs or trees. Distribution in PILA 100–1800 m, low elevation mixed forest, mixed forest in transition

Schradera obtusifolia C.M.Taylor to low elevation forest, mixed forest in transition to Epiphytic shrubs. Distribution in PILA 300–1200 m, low cloud forest. Global distribution: Endemic to Costa Rica. elevation mixed forest, mixed forest in transition to low Records: D. Santamaría 5933 (BM, INB, MO, PMA). elevation forest. Global distribution: Costa Rica, Panama. Records: G. Herrera 3325 (INB, MO). ℮ Citrus reticulata Blanco Mandarina (CR) .

ψ Sommera donnellsmithii Standl. Shrubs. Distribution in PILA 200–1200 m, low Mosqueta (CR). elevation mixed forest, mixed forest in transition to low Terrestrial shrubs or trees. Distribution in PILA elevation forest. Global distribution: Native to East Asia, 100–2100 m, low elevation mixed forest, mixed forest naturalised elsewhere. Records: D. Santamaría 3797 in transition to low elevation forest, mixed forest in (INB). transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Nicaragua to Colombia. Raputia heptaphylla Pittier Records : A.K. Monro 4803 (BM, INB), D. Solano 4343 Terrestrial shrubs or trees. Distribution in PILA 700–1500 (BM, INB, MO, PMA), D. Solano 4525 (BM, INB, MO, m, low elevation mixed forest, mixed forest in transition PMA). Regional conservation status: Near Threatened to low elevation forest, mixed forest in transition to (NT). cloud forest. Global distribution: Costa Rica to Ecuador, Venezuela, the Guyanas, Bolivia. Records: N. Zamora

Spermacoce capitata Ruiz & Pav. 4530 (BM, INB, MO). Terrestrial herbs. Distribution in PILA 100– 100 m, low elevation mixed forest. Global distribution: Belize to ψ Zanthoxylum melanostictum Schltdl. & Cham. Argentina. Records: D. Solano 2861 (INB), L. González Arcabú, Lagartillo, Lagarto colorado, Lagarto, Lagartillo 1199 (INB). colorado (CR). Terrestrial shrubs or trees. Distribution in PILA 1100–

Spermacoce ocymifolia Willd. ex Roem. & Schult. 3300 m, mixed forest in transition to low elevation forest,

mixed forest in transition to cloud forest, cloud forest, elevation forest, mixed forest in transition to cloud forest, low elevation oak forest, páramo. Global distribution: cloud forest, pasture, dense thicket. Global distribution: Mexico to Panama, Venezuela. Records: A. Chacón 485 Mexico, Honduras to Bolivia. Records: A. Rodríguez (INB), A.K. Monro 4098 (BM, INB, MO), B.B. Klitgaard 11578 (BM, INB, PMA), D. Santamaría 5946 (BM, 830 (BM, INB), B.B. Klitgaard 831 (BM, INB, MO), B. INB, PMA), D. Santamaría 6680 (BM, INB, PMA), D. Gamboa 588 (INB, MO), D. Solano 5088 (BM, INB, Santamaría 6731 (BM, INB, PMA), D. Santamaría 6765 MO, PMA), D. Solano 5139 (BM, INB, MO, PMA), E. (BM, INB, PMA), D. Santamaría 7054 (BM, INB, PMA), Alfaro 1321 (INB), E. Alfaro 3249 (INB), G. Davidse D. Solano 4832 (BM, INB, PMA).

25505 (MO), G. Davidse 28561 (CR, MO), G. Herrera

3683 (INB, MO), G. Herrera 5403 (CR, INB, MO), G. ψ Meliosma grandiflora C.V.Morton ex A.H.Gentry Herrera 6197 (CR, INB, MO), L. Acosta 2373 (INB), L. Terrestrial trees. Distribution in PILA 0–1800 m, low Angulo 202 (INB), L. González 3611 (INB), N. Zamora elevation mixed forest, mixed forest in transition to low 4130 (BM, INB, MO, PMA), N. Zamora 4167 (BM, INB, elevation forest, mixed forest in transition to cloud forest, MO, PMA), S. Knapp 10006 (BM, INB, MO), S. Knapp cloud forest. Global distribution: Costa Rica, Panama. 9859 (BM, INB, MO), S. Knapp 9901 (BM, INB, MO), Records: E. Alfaro 3704 (INB). Global conservation S. Knapp 9953 (BM, INB, MO). Regional conservation status: Vulnerable (VU), A2c.

status: Near Threatened (NT).

Meliosma idiopoda S.F.Blake

Terrestrial trees. Distribution in PILA 800–2500 m,

SABIACEAE mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low

Meliosma brenesii Standl. elevation oak forest, dense thicket. Global distribution: Terrestrial shrubs or trees. Distribution in PILA 100–1700 Mexico, Guatemala, El Salvador to Panama. Records: A. m, low elevation mixed forest, mixed forest in transition Rodríguez 11830 (BM, INB, PMA), D. Santamaría 7053 to low elevation forest, mixed forest in transition to cloud (BM, INB, PMA), D. Santamaría 7217 (BM, INB,PMA), forest. Global distribution: Costa Rica, Panama. Records: D. Santamaría 7232 (BM, INB, PMA), E. Alfaro 2197 A. Chacón 189 (INB), D. Solano 5451 (MO), D. Solano (INB), F. González 469 (INB), F. González 481 (INB), G. 5736 (MO), E. Alfaro 740 (INB). Davidse 28388 (CR, MO), G. Davidse 28411 (CR, MO).