Bombus (Pyrobombus) bimaculatus Cresson, 1863

Sheffield, Cory & Palmier, Kirsten M., 2023, Range expansion of Bombus (Pyrobombus) bimaculatus Cresson in Canada (Hymenoptera, Apidae), Biodiversity Data Journal 11, pp. 104657-104657 : 104657

Bombus (Pyrobombus) bimaculatus Cresson, 1863

Bombus Pyrobombus bimaculatus Bombus bimaculatus Cresson, 1863 - Cresson (1863): 92 [♂]. Holotype ♂. USA, Connecticut, by E. Norton [ANSP no. 2715].

Bombus Pyrobombus bimaculatus Bombus ridingsii Cresson, 1878 - Cresson (1878): 182 [♀]. Synonymy by Burks (1951): 1251, as a variety; synonymy by Mitchell (1962): 531. Syntypes ♀♀. USA, West Virginia, by J. Ridings [ANSP no. 2635].

Bombus Pyrobombus bimaculatus Bremus bimaculatus var. ahenus Bequaert and Plath, 1925 - Bequaert and Plath (1925): 275 [♀,♂]. Synonymy by Hurd (1979): 2198. Holotype ♀. USA, Massachusetts, Forest Hills, by O.E. Plath [MCZ no. 15,281].

Bombus Pyrobombus bimaculatus Bremus bimaculatus var. arboreti Bequaert and Plath, 1925 - Bequaert and Plath (1925): 276 [♀]. Synonymy by Hurd (1979): 2198. Holotype ♀. USA, Massachusetts, Forest Hills, on 2 June 1924, by O.E. Plath [MCZ no. 15,282].

Materials

Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 13835405 ; occurrenceRemarks: Research Grade; recordedBy: djsheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/20417139/medium.jpg?1530122554; occurrenceID: 3AD 960DF-5B0E-50D8-A677-9599386519DD; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Wascana Centre , Regina, SK, CA ; decimalLatitude: 50.4395783; decimalLongitude: -104.6165362; coordinateUncertaintyInMeters: 5; Event : samplingProtocol: none specified; eventDate: 06/27/2018; Record Level: source: https://www.inaturalist.org/observations/ 13835405 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 14775836 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/22048949/medium.jpg?1532664693; occurrenceID: 9E2AAA8C-90A5-585E-A31A-43B6BDE39E7A; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Victoria Ave , Regina, SK, CA ; decimalLatitude: 50.44732833; decimalLongitude: -104.6147217; coordinateUncertaintyInMeters: 10; Event : samplingProtocol: none specified; eventDate: 07/26/2018; Record Level: source: https://www.inaturalist.org/observations/ 14775836 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 14984878 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/22352937/medium.jpg?1533046158; occurrenceID: 8961341F-9246-536D-B163-7EB7CE1CC7A0; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Wascana Centre , Regina, SK, CA ; decimalLatitude: 50.43961333; decimalLongitude: -104.6165383; coordinateUncertaintyInMeters: 5; Event : samplingProtocol: none specified; eventDate: 07/29/2018; Record Level: source: https://www.inaturalist.org/observations/ 14984878 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 17006931 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/25717415/medium.jpg?1538165406; occurrenceID: C033564E-4784-5FFC-8601-D9D4C3C5C3C3; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Victoria Ave , Regina, SK, CA ; decimalLatitude: 50.44732833; decimalLongitude: -104.6147217; coordinateUncertaintyInMeters: 10; Event : samplingProtocol: none specified; eventDate: 07/26/2018; Record Level: source: https://www.inaturalist.org/observations/ 17006931 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 17006945 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/25717437/medium.jpg?1538165440; occurrenceID: 7ACD218F-B713-5CF4-968E-0AF2F4C76847; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Retallack St , Regina, SK, CA ; decimalLatitude: 50.44202833; decimalLongitude: -104.6226967; coordinateUncertaintyInMeters: 5; Event : samplingProtocol: none specified; eventDate: 07/28/2018; Record Level: source: https://www.inaturalist.org/observations/ 17006945 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 17006989 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/25717504/medium.jpg?1538165501; occurrenceID: 4467D420-E26D-5199-9811-48E 2AD 75E598; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Victoria Ave , Regina, SK, CA ; decimalLatitude: 50.44744167; decimalLongitude: -104.6139667; coordinateUncertaintyInMeters: 10; Event : samplingProtocol: none specified; eventDate: 07/26/2018; Record Level: source: https://www.inaturalist.org/observations/ 17006989 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 25003160 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/38686785/medium.jpg?1557678418; occurrenceID: 29EEC42D-273F-5E08-950C-9AA43F328DDE; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Retallack St , Regina, SK, CA ; decimalLatitude: 50.4419844; decimalLongitude: -104.6227672; coordinateUncertaintyInMeters: 22; Event : samplingProtocol: none specified; eventDate: 05/12/2019; Record Level: source: https://www.inaturalist.org/observations/ 25003160 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 25679542 ; occurrenceRemarks: Research Grade; recordedBy: sarah_306; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/39801310/medium.jpeg?1558714628; occurrenceID: 67452284-EDBD-5AA0-B315-F0F8E76C3A66; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Sherwood Dr @ Edward St (EB), Regina, CA ; decimalLatitude: 50.47503129; decimalLongitude: -104.6429203; coordinateUncertaintyInMeters: 9; georeferenceSources: gps; Event : samplingProtocol: none specified; eventDate: 05/23/2019; Record Level: source: https://www.inaturalist.org/observations/ 25679542 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 27533496 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/42848942/medium.jpg?1561328866; occurrenceID: F5773232-308B-5150-950B-17BF040C12EB; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Cameron St , Regina, SK, CA ; decimalLatitude: 50.43555833; decimalLongitude: -104.6255633; coordinateUncertaintyInMeters: 5; Event : samplingProtocol: none specified; eventDate: 06/23/2019; Record Level: source: https://www.inaturalist.org/observations/ 27533496 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 28411439 ; occurrenceRemarks: Research Grade; recordedBy: meghanmickelson; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/44288668/medium.jpeg?1562554211; occurrenceID: C2BFC522-0A1B-5C03-AF39-B27B612813D4; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Buena Vista, SK, CA ; decimalLatitude: 52.10832152; decimalLongitude: -106.6688233; coordinateUncertaintyInMeters: 55; Event : samplingProtocol: none specified; eventDate: 07/07/2019; Record Level: source: https://www.inaturalist.org/observations/ 28411439 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 29798922 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/46524789/medium.jpg?1564423555; occurrenceID: 1943D767-8FA1-5132-BAFC-B21E50038135; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: 15th Ave , Regina, SK, CA ; decimalLatitude: 50.44233; decimalLongitude: -104.6205833; coordinateUncertaintyInMeters: 10; Event : samplingProtocol: none specified; eventDate: 07/29/2019; Record Level: source: https://www.inaturalist.org/observations/ 29798922 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 29802698 ; occurrenceRemarks: Research Grade; recordedBy: djsheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/46530766/medium.jpg?1564426665; occurrenceID: 46EEAE46-E777-5CA1-A497-8B91D4CD2451; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: 15th Ave , Regina, SK, CA ; decimalLatitude: 50.44228183; decimalLongitude: -104.6206921; coordinateUncertaintyInMeters: 12; Event : samplingProtocol: none specified; eventDate: 07/29/2019; Record Level: source: https://www.inaturalist.org/observations/ 29802698 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 29806723 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/46537382/medium.jpeg?1564429997; occurrenceID: 23AB240B-EFC4-5ED7-86DF-FB4059F57D0C; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Cathedral , Regina, SK, Canada ; decimalLatitude: 50.44418732; decimalLongitude: -104.6196801; coordinateUncertaintyInMeters: 122; Event : samplingProtocol: none specified; eventDate: 07/29/2019; Record Level: source: https://www.inaturalist.org/observations/ 29806723 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 30221212 ; occurrenceRemarks: Research Grade; recordedBy: sarah_306; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/47204264/medium.jpeg?1564974488; occurrenceID: 375DD05A-DDC2-5B50-82C4-BF9A1268E32A; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Regina, SK, Canada ; decimalLatitude: 50.47494507; decimalLongitude: -104.6425247; georeferenceSources: gps; Event : samplingProtocol: none specified; eventDate: 08/04/2019; Record Level: source: https://www.inaturalist.org/observations/ 30221212 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 30221964 ; occurrenceRemarks: Research Grade; recordedBy: latebloomer2; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/47205538/medium.jpg?1564975375; occurrenceID: 40D0A612-FB4A-5BDC-8F7C-7951CF04C3D5; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Kiwanis Waterfall Park , Regina, SK, CA ; decimalLatitude: 50.43558528; decimalLongitude: -104.6219462; coordinateUncertaintyInMeters: 1273; Event : samplingProtocol: none specified; eventDate: 08/04/2019; Record Level: source: https://www.inaturalist.org/observations/ 30221964 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 30584482 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/47789579/medium.jpg?1565488686; occurrenceID: BA9FAE23-C0F8-5D00-9472-1ED212505ADB; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Retallack St , Regina, SK, CA ; decimalLatitude: 50.44203; decimalLongitude: -104.623; coordinateUncertaintyInMeters: 5; Event : samplingProtocol: none specified; eventDate: 08/10/2019; Record Level: source: https://www.inaturalist.org/observations/ 30584482 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 30880573 ; occurrenceRemarks: Research Grade; recordedBy: djsheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/48274049/medium.jpg?1565887254; occurrenceID: E47B0996-A460-5715-AD8A-08CC737FFB93; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Retallack St , Regina, SK, CA ; decimalLatitude: 50.44199455; decimalLongitude: -104.6229456; coordinateUncertaintyInMeters: 22; Event : samplingProtocol: none specified; eventDate: 08/15/2019; Record Level: source: https://www.inaturalist.org/observations/ 30880573 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 30940499 ; occurrenceRemarks: Research Grade; recordedBy: sarah_306; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/48373636/medium.jpeg?1565974844; occurrenceID: A85B689F-55BB-562B-9DC3-AAA115907FEA; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Sherwood Dr @ Edward St (EB), Regina, SK, CA ; decimalLatitude: 50.4749527; decimalLongitude: -104.6425705; georeferenceSources: gps; Event : samplingProtocol: none specified; eventDate: 08/16/2019; Record Level: source: https://www.inaturalist.org/observations/ 30940499 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 46539545 ; occurrenceRemarks: Research Grade; recordedBy: cwf_erin; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/73813750/medium.jpeg?1589928189; occurrenceID: 76853144-ED68-5D71-A769-2F6CF2895C32; Taxon: scientificName: Bombus bimaculatus; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Hymenoptera; family: Apidae; genus: Bombus; subgenus: Pyrobombus; Location: country: Canada; stateProvince: Saskatchewan; locality: Argyle St N, Regina, SK, CA ; decimalLatitude: 50.4806438; decimalLongitude: -104.6316069; coordinateUncertaintyInMeters: 161; Event: samplingProtocol: none specified; eventDate: 05/14/2020; Record Level: source: https://www.inaturalist.org/observations/ 46539545 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 46786006 ; occurrenceRemarks: Research Grade; recordedBy: kpalmier; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/74198671/medium.jpg?1590104851; occurrenceID: 8216655F-ABC7-5433-B536-D6F0FC882093; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Montague St , Regina, SK, CA ; decimalLatitude: 50.44485398; decimalLongitude: -104.6302722; coordinateUncertaintyInMeters: 4; Event : samplingProtocol: none specified; eventDate: 05/21/2020; Record Level: source: https://www.inaturalist.org/observations/ 46786006 Type status: Other material. Occurrence: occurrenceDetails: iNaturalist observation; catalogNumber: 51695891 ; occurrenceRemarks: Research Grade; recordedBy: cory_silas_sheffield; individualCount: 1; lifeStage: adult; associatedMedia: https://static.inaturalist.org/photos/82150777/medium.jpg?1593701998; occurrenceID: 1A6DBB21-AD3A-50CC-B0A3-DB0EB0A2BF15; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Wascana Creek , Regina, SK, CA ; decimalLatitude: 50.43746667; decimalLongitude: -104.6281283; coordinateUncertaintyInMeters: 6; Event : samplingProtocol: none specified; eventDate: 07/02/2020; Record Level: source: https://www.inaturalist.org/observations/ 51695891 Type status: Other material. Occurrence: catalogNumber: RSKM_ENT_E-226173 ; recordedBy: Ryan Oram; individualCount: 1; lifeStage: adult; occurrenceID: 87A39022-0660-5129-A358-DCF22F96445F; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Avonlea, SK, CA ; decimalLatitude: 50.0246; decimalLongitude: -104.9831; Event : samplingProtocol: netting; eventDate: 06/18/ 2019 Type status: Other material. Occurrence: catalogNumber: RSKM_ENT_E-189556 ; recordedBy: Kirsten Palmier; individualCount: 1; lifeStage: adult; occurrenceID: 88EB1216-984F-5531-A06B-2A5D4BE3D3EA; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Estevan, SK, CA ; decimalLatitude: 49.1475; decimalLongitude: -102.9673; Event : samplingProtocol: Blue vane trap; startDayOfYear: 07/05/2018; endDayOfYear: 08/03/ 2018 Type status: Other material. Occurrence: catalogNumber: RSKM_ENT_E-217788 ; recordedBy: Kirsten Palmier; individualCount: 1; lifeStage: adult; occurrenceID: 780F0200-5D87-5A96-B048-5D06D7466B98; Taxon : scientificName: Bombus bimaculatus; kingdom: Animalia ; phylum: Arthropoda ; class: Insecta ; order: Hymenoptera ; family: Apidae ; genus: Bombus ; subgenus: Pyrobombus ; Location : country: Canada; stateProvince: Saskatchewan; locality: Assiniboia, SK, CA ; decimalLatitude: 49.6343; decimalLongitude: -105.9931; Event : samplingProtocol: Blue vane trap; startDayOfYear: 07/27/2018 ; endDayOfYear: 08/14/2018

Distribution

Bombus bimaculatus is primarily a species of the eastern and central United States and Ontario and Quebec in Canada ( Colla et al. 2011, Williams et al. 2014, Koch et al. 2015) (Fig. 1 a, b Fig. 1 c Fig. 1 c). Since the publication of the most recent taxonomic treatment for North America ( Williams et al. 2014), its range has spread into the Maritime Provinces (New Brunswick, Nova Scotia, Prince Edward Island) of Canada and northwestwards into southern Saskatchewan (Fig. 1 c) and the adjacent United States.