Salcedia putzeysi ( Oberthuer , 1883)

Balkenohl, Michael, 2020, A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini), ZooKeys 901, pp. 1-81 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.901.39432
publication LSID	lsid:zoobank.org:pub:843F6065-D00E-48E6-9A26-9CEB044BF102
persistent identifier	https://treatment.plazi.org/id/E968F7C0-41BC-5F2A-81ED-DFC752C2DDB1
treatment provided by	ZooKeys by Pensoft
scientific name	Salcedia putzeysi ( Oberthuer , 1883)
status

Treatment

Salcedia putzeysi ( Oberthuer, 1883) View in CoL Figs 16 View Figures 14–19 , 34 View Figures 26–35 , 60 View Figures 58–63 , 75 View Figures 68–76 , 84 View Figure 84

Holoprizus putzeysi Oberthür, 1883: XL; Csiki 1927: 553.

Salcedia putzeysi ( Oberthür, 1883): Reichardt 1975: 103; Dostal 1993: 121; Lorenz 2005: 155.

Type material.

Holotype: ♂, with labels and data: beige, black framed and printed "Addah.W.Afrika Goldküste.” / white, handwritten "Holoprizus Putzeysi Oberth." / red, black printed “TYPE” (MNHN).

Remark.

The holotype was in two pieces but glued. The aedeagus was dissected and stored in a microvial at the same pin as the specimen. The specimen is complete. The type label was hidden under the determination label.

Additional material.

4 ♂, 1 ♀, 2 specs., with labels and data: blue, printed, black framed "MUSEUM PARIS DAHOMEY ENV. DE PORTO-NOVO WATERLOT 1912" (MNHN, CBB), 1 spec. with label: beige, handwritten " Salcedia schoutedeni All." (MNHN); 1 ♂, 1 ♀, with labels and data: white black printed "GHANA: Ashanti region Kumasi, Nhisau 330 m, N 6 43 - W1 36 Dr. S. ENDRODY-YOUNGA" / "Nr.284 black light 27.X.1967" (CBP).

Diagnosis.

A large sized species, with sub-elongate outline of the elytra with maximum width at middle and the pronotum with a short outer lateral carina near base. The pseudohumerus is rectangular and with an indistinct tooth. The antennomeres are elongate. Distinguished most clearly from the most similar species, S. schoutedeni and S. nigeriensis , by the head with an obtuse angle laterally at the base, the lateral outline of the elytra which are straight at middle, the less convex lateral margin of the pronotum, and by the different aedeagus. It differs most from S. matsumotoi sp. nov. by the different characters of the ratios of the elytra and the pronotum and the less numerous tubercles at the lateral margin.

Redescription.

Measurements in Table 1 View Table 1 .

Colour and surface: Griseous, shiny; top of carinae on head, pronotum and elytra as well as margins of pronotum and elytra opaque, covered with pale grey pili; legs and mandibles fuscous, antennae and palpi leoninous.

Head: Three-quarters of pronotum width. Outline campanulate. Clypeus wide, slightly convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by obtuse notches, with raised transverse field at middle, separated from frons by broad transverse furrow, frons with two raised paramedian carinae, joining anteriorly into a V-like keel, keel slightly tooth-like overhanging anteriorly, with two small glossy teeth bilaterally anterior to central keel, with a short distinctly diverging carina at each side paralaterally at base. Frons separated from supraorbital plates by flattened broad furrows; with conspicuously deep, broad pit at level of front of eyes; supra-antennal and supraorbital plates margined, margin of supraorbital plate distinctly raised, carina-like, supra-antennal plates slightly vaulted. Basal border with broad emargination at middle, obtuse angled laterally (angle 120-123°). Eyes large, convex, genae slightly convex, parts of both of them clearly visible from above, with slightly trapezoid shape in lateral view. Antenna with segments five to ten elongate (L/W 1.25), densely pubescent, segments two to four scarcely pubescent, scapus with sub-elongate reticulation. Labrum convex anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Mentum small, convex anteriorly, without tooth, epilobes wide, projecting and angled anteriorly, slightly margined medially, surface smooth, with isodiametric reticulation.

Pronotum (Fig. 34 View Figures 26–35 ): Outline rectangular, transverse, a third wider than long. Lateral margin slightly and regularly convex, maximum width at middle. Lateral margin distinctly crenulate, with ten to eleven distinct tubercles, tubercle anterior basal angle slightly more prominent, with two notches at posterior angles. Base straight laterally, with declining obtuse keel at middle pointing posteriorly. Disc flattened in lateral view, with two distinctly raised paramedian carinae parallel to median line and diverging posteriorly, without gap-like notches, with median line long, broadened anteriorly and narrowed posteriorly, with two additional and shortened carinae bilaterally at base, joining with the paramedian carinae and forming tooth-like tubercle at base pointing posteriorly, without anterior extension, without inner lateral carina, with very small and indistinct outer lateral carinae. All carinae sub-crenulate. Lateral margin broadly wing-like bent up, with six large and deep transverse pits, the basal ones partly separated into two smaller pits. Space between carinae and pits smooth.

Elytron: Slightly depressed in anterior half (lateral view), convex in frontal view. Sub-elongate, convex laterally but with straight part at middle, not diverging, maximum width at middle. Pseudohumerus rectangular, without distinctly projecting tooth. Apex rounded, with small but acute tooth at suture, slightly retracted. Disc with interneur six sub-crenulate, interneur one indistinctly carinate in anterior three-thirds, interneur three slightly carinate in whole length, interneur two running up to apex as nearly straight line, conspicuously raised, not reaching apex; interneur four running in parallel to interneur six, reaching base, shortened at apex. Interneur five and six with two rows of serial pits towards apex.

Hind wings: Fully developed.

Lower surface: Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with a row of pits, pits transverse towards apex, lateral margin of elytron smooth. Metepisternum elongate, with broad flat longitudinal groove. Metasternum, abdominal sternites one and five with numerous irregularly situated larger and smaller pits, sternite two to three smooth. Last abdominal sternites laterally with isodiametric reticulation, sternite one with longitudinal reticulation, two with isodiametric reticulation. Sternum four to six slightly sulcate. Sternum six with longitudinal flat pits laterally, with keel at apex.

Legs: Profemora with surface indistinctly reticulated. Protibia with robust, moderately curved terminal spine, laterally with four teeth of decreasing size and with equal distance from each other, dorsally and ventrally with two carinae. First tarsomeres distinctly elongated, as long as tarsomeres two to four together.

External sexual dimorphism: Not observed.

Male genitalia (Fig. 60 View Figures 58–63 ): Median lobe stout, in dorsal view distinctly arcuate, slightly cracked at middle and at beginning of apical spatula, in lateral view convex, with very few fine pili in apical half, apical part oblique, in cross section suboval at apex. Oroficium small. Endophallus with dense group of brush-like microtrichia. Dorsal paramere relatively long, bisinuate, with moderate apophyses; ventral one relatively long, shaped club-like; both parameres slightly distorted.

Female genitalia (Fig. 75 View Figures 68–76 ): Coxostylus slender, regularly broadened to base, less distinct curved, acute at apex, with eight large nematiform setae in basal half, one of them situated towards middle, one strong one at end of basal third.

Variation: The lateral margin of the pronotum varies from nearly straight (Ghana, Kumasi) to slightly more convex in specimens from Porto-Novo. The two specimens from the south of Ghana are slightly larger.

Distribution

(Fig. 84 View Figure 84 ). Known from a lagoon in Ivory Coast and from Benin as well as from the south of Ghana.

The specimens from Porto-Novo, Benin (formerly Dahomey) are quoted in Alluaud (1935) under “localités nouvelles" as S. schoutedeni .