Blakeius Ashmead, 1903

Turrisi, Giuseppe Fabrizio, Palmerini, Maurizio Matteini & Brothers, Denis J., 2015, Systematic revision and phylogeny of the genera Blakeius Ashmead, 1903 and Liomutilla André, 1907, with description of two new genera (Hymenoptera: Mutillidae, Myrmillinae), Zootaxa 4010 (1), pp. 1-78 : 9-10

publication ID	https://doi.org/ 10.11646/zootaxa.4010.1.1
publication LSID	lsid:zoobank.org:pub:E84D705A-B91B-49DA-9E23-F522C8614570
DOI	https://doi.org/10.5281/zenodo.5617580
persistent identifier	https://treatment.plazi.org/id/E93487B0-E757-FFA6-FF61-FA21FD126ED8
treatment provided by	Plazi
scientific name	Blakeius Ashmead, 1903
status

Treatment

Blakeius Ashmead, 1903: 327 .

Bisigilla Skorikov, 1927: 35 (as subgenus of Myrmilla Wesmael, 1851 ), type species Mutilla bipunctata Latreille, 1792 , ♀, by subsequent designation of Invrea 1964: 71. Junior subjective synonym of Blakeius Ashmead, 1903 according to Brothers 1975: 591.

Type species. Mutilla bituberculata Smith, 1855 (♀), by original designation, junior subjective synonym of Mutilla bipunctata Latreille, 1792 (♀) according to Morawitz (1865): 125 ( Lelej & Brothers 2008).

Diagnosis. ♀. Supra-antennal tooth present, acute projection produced by frontal carina, more or less medially expanded; frontal carina touching inner margin of eye; antennal rim mostly concealed within the frons, not visible dorsally; gena (lateral view) convex, bulging, not angulate, without tooth-like process; T2 with two very wide circular whitish to slightly goldish setal spots disposed transversely, with setae very dense and recumbent; spots close to anterior margin of T2.

♂. Head strongly convergent toward occipital margin; longitudinal frontal furrow absent; ocelli well developed; notaulus and parapsidal line absent; tegula and wings well developed, not reduced; T2 without pale spots; S8 with sides subparallel bearing single raised medial tubercle; genital capsule with cuspis of volsella elongate, finger-shaped, apex regularly rounded, weakly convex to flat toward apex, ending well beyond mid length of paramere harpe, without longitudinal carina; digitus of volsella much shorter than cuspis, about half its length; midventral process of penis valve present, elongate and strongly curved.

Redescription. ♀. Head mostly blackish, sometimes lighter with antennal rim concolorous with rest of head ( Figs 3 View FIGURE 3 A, 3B, 3C); mesosoma reddish orange, with lateral and ventral parts extensively dark brown to blackish ( Fig. 2 View FIGURE 2 B); metasoma entirely blackish; integument of body coarsely sculptured, dull; head regularly rounded behind eyes, with dense and coarse punctures, without whitish setal spot on vertex and occiput ( Fig. 3 View FIGURE 3 A); supraantennal tooth present as acute projection produced by frontal carina, more or less expanded medially ( Fig. 3 View FIGURE 3 A); antennal rim mostly concealed within frons, not visible dorsally ( Figs 3 View FIGURE 3 A, 3C); A1 with ventro-lateral carina; A3 more than twice length of A2; frontal carina touching inner margin of eye; postgena (behind oral fossa) convex ( Fig. 4 View FIGURE 4 A) or narrowly and deeply concave behind elevated tuberculate hypostomal carina, bowl-shaped ( Fig. 12 View FIGURE 12 A); occipital margin straight; occipital carina a narrow rim ( Fig. 4 View FIGURE 4 A); gena (lateral view) convex, bulging, not angulate, without tooth-like process ( Fig. 7 View FIGURE 7 B); genal carina absent; clypeus not prominent, wider than high, anterior margin wide, slightly concave with lateral tooth-like projection ( Fig. 3 View FIGURE 3 E); oral fossa wider than long, roughly V-shaped ( Fig. 4 View FIGURE 4 C); mandible with inner apical tooth normally shaped, not curved ( Fig. 3 View FIGURE 3 D); mandible with inner basal tooth ( Figs 3 View FIGURE 3 C, 3D); pronotum about as wide as propodeum ( Fig. 2 View FIGURE 2 A); pronotum with epomial carina well developed, close to pronotal spiracle; fore coxa without distal tooth-like process; mid tibia with 6–7 dorsal pre-apical spines arranged in two rows; hind tibia with 5–7 dorsal pre-apical spines arranged in two rows; T2 with two sharply defined whitish to goldish setal spots disposed transversely, close to anterior margin ( Figs 2 View FIGURE 2 A, 2B); T3 with broad pubescent band, entire or very weakly but distinctly interrupted medially; T4 without ( Figs 2 View FIGURE 2 A, 2B) or with broad pubescent band, very weakly but distinctly interrupted medially ( Figs 6 View FIGURE 6 A, 6B); S6 flat or slightly folded longitudinally.

♂. Head mostly blackish, sometimes lighter, with antennal rim concolorous with rest of head ( Figs 3 View FIGURE 3 F, 3G, 3H); mesosoma reddish orange, with lateral and ventral parts extensively dark brown to blackish ( Fig. 2 View FIGURE 2 D); metasoma entirely blackish; integument of body coarsely sculptured, dull; head strongly convergent posteriorly ( Fig. 3 View FIGURE 3 F), with dense and coarse punctures; A1 with ventro-lateral carina; A3 shorter than A4; ocelli well developed ( Fig. 7 View FIGURE 7 F); longitudinal frontal furrow absent ( Fig. 7 View FIGURE 7 F); occipital carina (lower part) a narrow rim; clypeus not prominent, wider than high, anterior margin wide, moderately concave with lateral tooth-like projection ( Fig. 3 View FIGURE 3 J); oral fossa wider than long, roughly V-shaped; mandible with inner apical tooth straight, not curved ( Fig. 3 View FIGURE 3 I); pronotum with sides convergent toward anterior margin ( Fig. 2 View FIGURE 2 C); posterior margin of pronotum, V-shaped ( Fig. 8 View FIGURE 8 B); pronotum, median length less than half lateral length (pronotum short) ( Fig. 8 View FIGURE 8 B); mesoscutum convex ( Fig. 6 View FIGURE 6 D); notaulus and parapsidal line absent; transscutal suture well developed, evident ( Fig. 8 View FIGURE 8 B); tegula sub-circular, well developed, normal sized ( Fig. 8 View FIGURE 8 B); fore and hind wings present and well developed ( Figs 2 View FIGURE 2 C, 2D); fore wing, submarginal cell 2 similar in size to submarginal cell 1 ( Fig. 4 View FIGURE 4 E); fore wing with vein 2m-cu present ( Fig. 4 View FIGURE 4 E); mesoscutellum and metascutellum well defined ( Fig. 8 View FIGURE 8 B); anterior and posterior metanotal carinae well raised; propodeum (lateral view) sloping ( Fig. 6 View FIGURE 6 D), narrower than or as wide as pronotum (dorsal view) ( Fig. 8 View FIGURE 8 B); mesopleuron strongly convex, strongly rugose-punctate, width of upper part about half width of lower part; mid tibia with one dorsal pre-apical spine; hind tibia with one dorso-lateral pre-apical spine; T1 campanulate ( Figs 6 View FIGURE 6 C, 6D); T2 without setal or cuticular spots ( Figs 2 View FIGURE 2 C); T3 with broad pubescent band, entire or very weakly but distinctly interrupted medially ( Figs 2 View FIGURE 2 C, 6C); T4 with or without pubescent band, if present weakly interrupted medially ( Figs 6 View FIGURE 6 C, 6D); S2 with medial well developed, tooth-like process ( Fig. 2 View FIGURE 2 D); S8 with sides subparallel ( Fig. 5 View FIGURE 5 F), and well developed stout tooth-like median process ( Fig. 5 View FIGURE 5 G); genital capsule with cuspis of volsella elongate, finger-shaped, apex regularly rounded, weakly convex to flat toward apex, ending well beyond mid length of paramere harpe, without longitudinal carina ( Fig. 5 View FIGURE 5 B); digitus of volsella much shorter than cuspis, about half its length ( Fig. 5 View FIGURE 5 D); midventral process of penis valve present, elongate and strongly curved ( Fig. 5 View FIGURE 5 E).