Calendula suffruticosa subsp. guruguensis A.C. Gonç & P.Silveira, 2023
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publication ID |
https://doi.org/ 10.11646/phytotaxa.605.1.1 |
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persistent identifier |
https://treatment.plazi.org/id/E82AFC0A-1560-CA5E-66ED-FB3C820FFE35 |
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treatment provided by |
Plazi |
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scientific name |
Calendula suffruticosa subsp. guruguensis A.C. Gonç & P.Silveira |
| status |
subsp. nov. |
11f. Calendula suffruticosa subsp. guruguensis A.C. Gonç & P.Silveira , subsp . nov. Type:— MOROCCO.Al Hoceïma, Gurugu, 950 m, [35°19’ N, 2°57’ W], 19 May 1932, Sennen & Mauricio 8438 (holotype BM! [BM000813672]; isotypes BC! [139240], G!).
Diagnosis:— Calendula suffruticosa subsp. guruguensis differs from C. suffruticosa subsp. suffruticosa by the presence of trialate achenes.
It differs from C. suffruticosa subsp. marsaensis by having middle achenes trialate, (6.3) 8.5–9.8 (10.3) mm, with length/width approx. 1, instead of bi-trialate, (10.5) 13.6–16 (22) mm, with length/width> 1.5.
Description:––Perennial herbs. Stems (37) 44.5–75.5 (81) cm long, prostrate to decumbent, with glandular hairs distally and predominance of non-glandular white-arachnoid pubescence proximally, not viscid/viscous. Basal leaves (4.8) 5–7.9 (11.7) × (1.1) 1.5–2.1 (2.9) cm, 0.2–0.3 mm thick, oblanceolate to spatulate, apex obtuse, or sometimes ± acute, margins sub-entire to sinuate-dentate, with white-arachnoid pubescence. Capitula solitary (2.3) 3.1–3.9 (4.0) cm diameter. Outer achenes rostrate (12) 16.5–25 × (1.7) 2.2–2.4 (2.5) mm, straight or slightly curved, without dorsal spines, sometimes with 1–2 teeth, at the base or at the apex; middle achenes trialate (6.3) 8.4–9.8 (10.3) × (5.3) 6–9.1 (10) mm, with a rostrum 2–3.6 (3.7) mm; inner vermiculate-exalate 4–4.3 × 1.7–1.9 mm, hemicyclic-falcate. Figure 17 View FIGURE 17 .
Habitat and distribution:—As far as we know, this taxon grows on greywacke rocks, in semi-arid Mediterranean climate, at elevations from sea level to probably 800 m, in mounts Gurugu and Kebdana, near Mellila, and near sea level at Al Hoceïma.
Conservation status:—Critically Endangered (CR). The taxon is under numerous threats, especially due to climate change (extreme weather events) and drought. The species is rare and local with a small-restricted range. In the single population known by th team, the number of mature individuals is estimated to be <10. The AOO is 4 km 2, which qualifies for CR. Considering the number of individuals, the reduced size of the population and of its habitat, which puts it under a high risk of extinction due to climate changes, or other unexpected threats, we propose a conservation status of CR B2ab(ii, iv, v).
Chromosome number:— 2n = 32. Genome size:—2.96 ± 0.05 pg. Eponymy :— For Gurugu Mount in Morocco where it occurs. Additional collections:— MOROCCO. Melilla : Melilla, Barranco del Nano, 1 June 1915, A . Caballero s.n. ( MA!) ;
Melilla, Pico de la Peineta, 13 April 1943, A. Arnal s.n. ( MA!); Jbel Gurugú, 562 m [35°14’11” N, 2°58’29” W], 11 GoogleMaps
June 2012, P. Silveira & A.C.R.S. Gonçalves 3265 (AVE!). Tanger-Tetouan-Al Hoceïma: Nador, Al Hoceima, cabo ao
SE, junto a urbanización, [35°14’36” N, 3°55’51” W], 11 June 1993, J. M. Montserrat, T. Abdelkader , J. Molero, J. Pallàs GoogleMaps ,
J. Vicens & M. Veny JMM 3585 5 (BC!).
| A |
Harvard University - Arnold Arboretum |
| MA |
Real Jardín Botánico |
| J |
University of the Witwatersrand |
| M |
Botanische Staatssammlung München |
| T |
Tavera, Department of Geology and Geophysics |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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