Stachys, Linnaeus, 1753

Akçiçek, Ekrem, 2020, Taxonomic revision of Stachys sect. Olisia (Lamiaceae: Lamioideae) in Turkey, Phytotaxa 449 (2), pp. 109-148 : 113-115

Stachys L. sect. Olisia Dumort., Fl. Belg. : 44 (1827) ( Figs. 1 , 2 )

Type:― S. annua (L.) L., Sp. Pl. ed. 2: 813 (1763)

Description:―Suffruticose perennials, rarely biennials to annuals, with or without basal rosettes. Flowering stems erect to decumbent, simple or branched. Basal leaves ovate to ovate-oblong or ovate-lanceolate, sometimes orbicular, cordate to cuneate or attenuate at base, passing either gradually or abruptly into cauline leaves. Cauline leaves ovate-lanceolate to lanceolate or oblong-lanceolate to oblanceolate, cuneate to attenuate at base, gradually passing into smaller sessile floral leaves. Bracteoles usually few, setaceous, 0.5‒3 mm, rarely herbaceous, lanceolate to linear-subulate, mucronate, rarely absent. Calyx usually subbilabiate, ± regular, tubular to campanulate, teeth subequal, triangular to lanceolate or lanceolate-subulate, mucronate. Corolla tube usually subexserted, annulate (except S. inanis ). Nutlets oblong-elliptic to obovoid, usually minutely reticulate, rarely smooth on surface, black or blackish-brown.

Distribution:―Mainly Mediterranean Europe and Anatolia, extending into Iran and Caucasia (24 species in the Old World area).

Notes:―The section Olisia of Stachys was described by Dumortier (1827), and later treated at generic rank by Spach (1838: 166). Bhattacharjee (1980) divided S. sect. Olisia into 5 subsections: Rectae R. Bhattacharjee (1980: 84), Distantes R. Bhattacharjee (1980: 86), Annuae R. Bhattacharjee (1980: 86), Rosulatae R. Bhattacharjee (1980: 87), and Spinosae R. Bhattacharjee (1980: 87) . To date, however, Bhattacharjee’s (1980) classification has not been well supported by molecular data ( Scheen et al 2010, Bendiksby et al. 2011, Salmaki et al. 2013, 2019). Therefore, the subsectional classification was not applied herein.

Figures

FIGURE 1. A. Stachys recta subsp. subcrenata (Ö. Güner 2412); B. S. atherocalyx (Akçiçek 5637); C. S. iberica subsp. iberica var. iberica (Akçiçek 5588); D. S. hakkariensis (M. Fırat 32600); E. S. sparsipilosa (Akçiçek 5598); F. S. aleurites (Akçiçek 5601); G. S. bombycina (Akçiçek 5550). Photograph D by M. Fırat.

FIGURE 2. A. Stachys annua subsp. annua (Akçiçek 5559); B. S. distans var. cilicica (Akçiçek 5527); C. S. diversifolia (Akçiçek 5595); D. S. inanis (Akçiçek 5630); E. S. maritima (Akçiçek 5624); F. S. distans var. distans (Akçiçek 5807); G. S. munzurdagensis (Akçiçek 5332).