Stachys angustifolia M.Bieb., Fl. Taur.

Akçiçek, Ekrem, 2020, Taxonomic revision of Stachys sect. Olisia (Lamiaceae: Lamioideae) in Turkey, Phytotaxa 449 (2), pp. 109-148 : 116-120

2. Stachys angustifolia M.Bieb., Fl. Taur. -Caucas. 2: 52 (1808) ( Figs. 5, 8 )

= S. tenuifolia Pall. ex Link, Enum. Hort. Berol. Alt. 2: 109 (1822), nom. illeg. = S. tenuifolia Janka, Oesterr. Bot. Z. 22: 180 (1872), nom. illeg. = S. abrotanifolia Boiss. & Heldr. in Boiss., Fl. Orient. 4: 732 (1879) = S. angustifolia var. haynaldii Janka ex Nyman, Consp. Fl. Eur. : 580 (1881), nom. nud. = S. angustifolia subsp. tenuifolia Nyman, Consp. Fl. Eur. : 580 (1881)

Type:—[Crimea] in Tauriae meridionalis petrosis sylvaticis, circa pagum Temirdschi, Marschall von Bieberstein s.n. (holo: LE).

Description:―Suffrutescent, green, perennial herbs without basal rosettes of leaves. Flowering stems 25‒65 cm, slender, numerous, erect, branched, glabrous, with few sessile to short glandular hairs. Lower and middle cauline leaves laciniate to pinnatipartite with linear or filiform segments, 25‒45 × 0.7‒2 mm, entire, passing into narrowly linear to linear-lanceolate, entire, upper cauline and floral leaves 1.5‒25 mm. Floral leaves (1.5‒) 2‒25 mm, longer or shorter than verticillasters, acute at apex, shortly spinescent tipped. Verticillasters (3‒)8‒15(‒19), remote, (0.5‒3.5 cm distant), 2- flowered. Bracteoles setaceous, minute, 0.5‒1.5(‒2) mm. Pedicels 2‒5 mm. Calyx ± regular, subcampanulate, inflated in fruit, 6‒7 mm, distinctly 10-nerved, glabrous with few sessile glands mixed with adpressed hairs, tubercled at base, mouth without a ring of hairs; teeth ± equal c. 2 mm, triangular, spinescent-tipped (spine 0.5 mm), ± half the length of tube, slightly recurved. Corolla yellow tinged with pink, 12‒16 mm, tube exserted, annulate; limb bilabiate, upper lip 3‒6 mm, entire, thinly covered with short adpressed hairs not exceeding lip outside, lower lip 3-lobed, middle lobe larger than 2 lateral lobes, entire, 4‒7 mm. Stamens 4, slightly exserted from corolla tube (less than half-way along upper corolla lip). Nutlets broadly obovoid, faintly trigonous, slightly winged near base, 1.8‒2 × 1.5 mm, minutely reticulate-foveate, blackish-brown.

Phenology: ―Flowering between June and July.

Distribution and ecology: ― Greece, Bulgaria, Romania, Crimea, South Russia, Turkey in Europe. Euro-Siberian element. Rocky places, stony slopes, hills, under Quercus , 10–500 m elevation.

Conservation status:―VU [Criteria B2ab(i,ii,iv)] ( IUCN 2017). It is known from less than 10 localities, and the estimated area of occupancy is smaller than 2000 km ².

Remarks: ― Stachys angustifolia M.Bieb. is most closely allied to S. iberica Marschall von Bieberstein (1808: 51) , from which it differs strikingly in having linear pinnatipartite cauline leaves, glabrescent stems, 2-flowered verticillasters, 2‒5 cm pedicels, distinctly 10-nerved calyx, triangular teeth, and yellow tinged and exserted corolla.

Figures

FIGURE 8. Distribution map of Stachys angustifolia (▲), S. atherocalyx (●), S. hakkariensis (▼), S. recta subsp. subcrenata (▃) and S. sparsipilosa (⊕) in Turkey.