Pupina tchehelensis Morgan, 1885

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina tchehelensis Morgan, 1885

Figs 18D , 37A-C , 39C, D

Pupina tchehelensis Morgan, 1885: 414, 415, pl. 7, fig. 4. Type locality: mont Tchéhèl [possibly the hill in the vicinity of Ipoh, Perak, Malaysia]. von Möllendorff 1891: 346, Bukit Pondong. Maassen 2001: 41. BEDO 2017: 94.

Pupina artata [non Benson]- von Möllendorff 1887 [1886]: 314. von Möllendorff 1891: 345, 346.

Pupina (Tylotoechus) tchehelensis - Kobelt 1902: 323. Laidlaw 1928: 34.

Pupina tchechelensis [sic]- van Benthem Jutting 1949: 57, Sungei Siput, Perak.

Material examined.

SMF 109947/6 (6 shells; Figs 37A , 39C ) from Bukit Pondong, Perak. CUMZ 12135 (1 shell; Fig. 37B ) from Tham Suea Temple, Mueang Krabi District, Krabi Province, 6 Oct. 2006. CUMZ 12136 (7 shells; Figs 18D , 37C , 39D ) from limestone mountain, Phang Nga Province, 1 May 1999.

Diagnosis.

Shell ovate; last whorl ca. 70-75% of shell height. Apertural lip slightly thickened but not expanded. Parietal tooth sharp, tooth-like, thickened; columellar tooth fin-shaped, slightly thickened, located next to slit-like anterior canal. Posterior canal gradually widening like a keyhole.

Differential diagnosis.

Pupina tchehelensis is most similar to P. lowi and P. brachysoma in having a sharp, tooth-like, thickened parietal tooth, a fin-shaped, thickened, columellar tooth that is located next to a slit-like anterior canal, and a posterior canal that is gradually widening. However, P. tchehelensis is different from P. lowi by having a more ovate shell shape, and differs from P. brachysoma in that the apertural lip is not expanded.

Distribution.

Malaysia ( Maassen 2001) and southern Thailand.

Remarks.

Both similar species P. tchehelensis and P. lowi were originally described by de Morgan (1885) from the same vicinity within Perak, peninsular Malaysia: de Morgan (1885) stated that P. lowi is "much larger than P. tchehelensis , and this species is distinguished by the shape of its whorls which are much more flattened." As the type materials of P. tchehelensis have not yet been discovered, and P. tchehelensis specimens have a slightly higher shell than P. lowi , we do not synonymise P. tchehelensis with P. lowi . Specimens from Thailand have a larger shell than those from Perak, Malaysia (Fig. 37A-C ).

Figures

Figure 18. External shell morphology of three mainland Southeast Asian Pupina species groups A Pupina artata from the Pupina artata species group, CUMZ 12003 from Ban Ping Khong, Chiang Mai B Pupina peguensis from the Pupina arula species group, CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai C Pupina siamensis from the Pupina arula species group, CUMZ 12052 from Sri Thong Cave, Sra Keo, and D Pupina tchehelensis from the Pupina aureola species group, CUMZ 12136 from limestone mountain, Phang Nga. Red frames focus on the parietal tooth and posterior canal; blue frames focus on the curvature of the apertural lip when observed from lateral view.

Figure 37. Shells of Pupina aureola species group from mainland Southeast Asia. A-C Pupina tchehelensis: specimens A SMF 109947 / 6 B CUMZ 12135, and C CUMZ 12136 D, E Pupina latisulci sp. nov. D holotype CUMZ 12146 / 1 and E paratype CUMZ 12146 / 2 F-I Pupina paviei F paralectotype MNHN-IM- 2000 - 35837 G paralectotype RBINS 525404, and specimens H CUMZ 12134 and I NHMUK ex. Dautzenberg coll. J Pupina tongupensis, syntype NHMUK 1906.4.4.38 K Pupina lowi, syntype MNHN-IM- 2000 - 35846 L Pupina anceyi, syntype MNHN-IM- 2000 - 35833 M Pupina laffonti, syntype MNHN-IM- 2000 - 9656 N Pupina solidula, lectotype SMF 109915 / 1 O Pupina douvillei, holotype MNHN-IM- 2000 - 35532. Photo: M. Caballer, P. Maestrati, MNHN (F, K-O), F. Trus, RBINS (G), J. Ablett, H. Taylor, NHM (J).

Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu.