Carabus (Lipaster) stjernvalli, Mannerheim, 1830

Fominykh, Dmitriy D., Zamotajlov, Alexandr S., Khomitskiy, Eugeniy E. & Titarenko, Andrey Yu., 2020, Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini), Zootaxa 4729 (3), pp. 301-328 : 302-303

1. Carabus (Lipaster) stjernvalli subsp.?

( Fig. 1 )

Material examined. 113♂♂, 94♀♀, Georgia, Samtskhe-Dzhavakheti , Gorelovka vill. env., SW slopes of Madatapa Mt. near Madatapa Lake, 2285 m, N 41°12’58.80” / E 43°47’46.60”, pitfall traps, 09.06.- 11.07.2018 (D. Fominykh) ( ZISP, MPSU, CFDD, CTAY, CZAM, CEEK).

Remarks. Surprisingly this population resembles more the spatially remote nominotypical subspecies than the neighboring geographic races ( Table 1 , Fig. 115 ). Probably it deserves status of a separate taxon, however we still do not have enough comparative material from adjacent territories for a more definite judgment.

Figures

FIGURE 1. Collecting sites of Carabus (Lipaster) stjernvalli Mannerheim in the South Caucasus: red square—C. stjernvalli ssp., Gorelovka env.; green square—C. stjernvalli paravanicus subsp. nov.; pink rhombs—C. stjernvalli tabackuriensis Novotný & Vořišek; red rhombus—C. stjernvalli tabackuriensis Novotný & Vořišek, type locality; red triangles—C. stjernvalli kartalinicus subsp. nov.; blue circle—C. stjernvalli bogatshevi Zamotajlov, type locality; violet empty squares—C. stjernvalli stjernvalli Mannerheim; red crosses—C. stjernvalli gvalijai Retezár.

FIGURE 115. Distribution of morphometric characters in some Carabus (Lipaster) stjernvalli Mannerheim subspecies constructed using discriminant analysis based on 8 parameters (see text for measurements details): red squares—C. stjernvalli ssp., Gorelovka env.; green squares—C. stjernvalli paravanicus subsp. nov.; violet empty squares—C. stjernvalli stjernvalli Mannerheim.