Pison hypostomale Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 222-224

Pison hypostomale Pulawski , species nova

Figures 500 -507.

NAME DERIVATION.– Hypostomale is a Neolatin neuter adjective derived from hypostoma; with reference to the unusually high hypostomal carina.

RECOGNITION. – Pison hypostomale is an all black species, with three submarginal cells, the second recurrent vein interstitial with the second intersubmarginal vein, and the setae appressed on tergum I. Like P. pilbara and P. separatum , it is characterized by an expanded hypostomal carina ( Fig. 502 ), about as wide as 0.5 × midocellar diameter and markedly wider than the occipital carina. Unlike P. pilbara (whose female is unknown), it has the scutal punctures less than one diameter apart and the scutal setae appressed (rather than punctures 2-3 diameters apart and the setae erect). It differs from P. separatum by a number of characters: the setae are appressed and oriented ventrally between the antennal socket and the midocellus ( Fig. 503 ) rather than oriented dorsolaterad in the frons dorsal half, the occipital carina is of the usual height (rather than expanded mesodorsally), the ocellocular distance is equal to about 0.7 × hindocellar diameter in females, about 1.1-1.3 × in males (rather than 0.3 × in females and 0.6-0.7 in males), the scutal and mesopleural interspaces are dull, conspicuously microsculptured (rather than shiny, unsculptured), the propodeal dorsum has conspicuous ridges joining the longitudinal carina that separates the dorsum from the side (no such ridges in P. separatum ), the clypeal lamella of the male is obtusely angulate ( Fig. 501 ) rather than acutely angulate, and male sternum VIII has the apical margin conabout one diameter apart below midpoint (mesopleural punctures less than one diameter apart in P. laterirugosum ).

DESCRIPTION.– Frons dull, minutely punctate, punctures barely recognizable, less than one diameter apart, middle supraantennal carina absent or evanescent. Hypostomal carina expanded ( Fig. 502 ), about as wide as 0.5 × midocellar diameter and markedly wider than occipital carina. Labrum not emarginate. Anteromedian pronotal pit transversely elongate, almost as long as 1.5 × midocellar diameter. Scutum slightly foveate along flange, with inconspicuous longitudinal ridges adjacent to posterior margin; scutal punctures minute, less than one diameter apart, interspaces dull, markedly microsculptured. Tegula somewhat enlarged. Mesopleural punctures well defined, averaging more than one diameter apart at center; interspaces dull, markedly microsculptured. Postspiracular carina present, about 1.5 × as long as midocellar diameter. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged (ridges becoming quite conspicuous adjacent to longitudinal carina); side ridged, punctate between ridges; posterior surface coarsely ridged. Posteroventral forefemoral surface with fine punctures about 2-3 diameters apart. Hindcoxal dorsum with outer margin not carinate. Punctures of tergum I about two diameters apart on horizontal part medially (compressed against each other on apical depression). Punctures of sternum II several diameters apart apicomesally, apical depression impunctate.

Setae silvery, appressed on scutum and tergum I, on frons strictly appressed, oriented ventrally ( Fig. 503 ), on lower gena subappressed, curved, up to about 1.3 × midocellar diameter long, partly concealing integument on clypeus. Apical depressions of terga with silvery setal fasciae.

Head, thorax, propodeum, legs, and gaster black, mandible black, brown apically.

♀.– Upper interocular distance equal to 0.52 × lower interocular distance; ocellocular distance equal to 0.7 × hindocellar diameter, distance between hindocelli equal to 0.9 × hindocellar diameter; eye height equal to 1.06 × distance between eye notches. Free margin of clypeal lamella broadly, obtusely angulate ( Fig. 500 ). Dorsal length of flagellomere I 2.4 × apical width, of flagellomere IX 1.4 × apical width. Mandible: trimmal carina with minute preapical tooth. Length 6.8 mm; head width 2.2 mm.

♂.– Upper interocular distance equal to 0.62-0.66 × lower interocular distance; ocellocular distance equal to 1.1-1.3 × hindocellar diameter, distance between hindocelli equal to 1.0-1.3 × hindocellar diameter; eye height equal to 1.06-1.10 × distance between eye notches. Free margin of clypeal lamella obtusely, broadly angulate ( Fig. 501 ). Dorsal length of flagellomere I 2.4-2.5 × apical width, of flagellomere X 1.4 × apical width. Sternum VIII punctate, its apical margin convex except concave laterally, with sharply angulate apicolateral angle ( Fig. 504 ). Genitalia: Figs. 505, 506 . Length 6.0- 6.4 mm; head width 1.9-2.0 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 507).–

Eastern Queensland.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA:

Queensland: Dalby, 30 Oct 1979, H.E. Evans, M.A.

Evans, and A. Hook (QMB, registration number

T228762).

PARATYPES: AUSTRALIA: Queensland: near

Brisbane Forest Park at 27°26.0ʹS 152°55.4ʹE,

FIGURE 507. Collecting localities of Pison hypostomale 19 Oct 2006, V. Ahrens and W.J. Pulawski (2 ♀, 3 ♂,

Pulawski, sp. nov.

CAS; 1 ♀, QMB ).

Figures

FIGURES 500-503. Pison hypostomale Pulawski, sp. nov. (500) Female clypeus and mandibles; (501) Male clypeus and mandibles; (502). Posterior surface of female head in oblique view (arrow shows hypostomal carina); (503) Male frons showing orientation of setae.

Figures 504-506. Pison hypostomale Pulawski, sp. nov., male. (504) Sternum VIII (ventral surface); (505) Genitalia in dorsal view; (506) Genitalia in lateral view.